That inheritance from both parents is only a secondary consideration
in regard to the increase of a part by artificial selection is made
evident by the fact that _many secondary sexual characters_ have been
modified, although the breeder selected only in regard to one parent.
Nevertheless in this very domain the greatest results have been
achieved; witness the Japanese breed of cocks with tail-feathers six
feet long. This astonishing result has been reached by the strictest
selection of the cocks in which the feathers were a little longer
than those of other cocks, and the increase in the length of feathers
depended--according to our theory--simply on the fact that, by the
selection of the determinants which were already varying in the
direction of increased length, this process of increase was guarded
from interruption by chance unfavourable conditions of nutrition.
The continuance of variation in the upward direction in which it had
already started is not effected directly by personal selection, but
is so indirectly, for without this constant fresh intervention of
selection the increase would be apt to come to a standstill, or the
variation might even take a contrary direction. There are two other
factors operative to which we have not yet given sufficient attention.
They are, the multiplicity of the ids in every germ-plasm, and sexual
reproduction.
If--as we must assume--each germ-plasm is made up of several or many
ids, there must be several or many determinants of each part of the
organism, for each id contains potentially the whole organism, though
with some individuality of expression. The child is thus not determined
by the determinants of a single id, but by those of many ids, and the
variations of any part of the body do not depend on the variations of a
single determinant _X_, but on the co-operation of all the determinants
_X_ which are contained in the collective ids of the relevant
germ-plasm. Thus it is only when a majority of the determinants have
varied upwards or downwards that they dominate collectively the
development of the part _X´_ and cause it to be larger or smaller.
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