There are even characters the biological use of which is unknown to
us, but in regard to which we can affirm that they have a use. Thus
Eigenmann described the larva of an American eel, which differs from
other so-called 'Leptocephali' in that a row of seven black spots runs
along its side. Apparently all these lie upon the side turned towards
us, but in reality they are distributed on both sides, three lying on
the left and four on the right, and so arranged that they look like a
single row of spots at regular intervals, for the flat little fish is
absolutely transparent. The habits of this larva are not yet known, but
we may conclude that this appearance of a simple row of spots must have
some value for the animal, for such a significant asymmetry could not
have arisen for purely internal reasons (Fig. 107, _C_). It is possible
that the fish is thus made to resemble parts of some marine alga, and
that it is thereby protected from many enemies; that there is not a
complete row upon each side may depend upon the fact that the two rows
would be visible at the same time, and that they would blur each other
in the eyes of the swimming enemy, and so destroy the resemblance of
the picture to its unknown model.
But it cannot be denied that there are characters which have no special
biological significance. There are doubtless many such characters,
which stand beyond the threshold of good or bad, and which are
therefore not affected by personal selection; it is difficult and often
impossible to point these out with certainty. The shape of the human
nose and of the human ear, the colour of the hair and of the iris, may
be such indifferent characters whose peculiarities are to be referred
solely to germinal selection. On the other hand, I would not venture to
assert that the gay colouring and the complex markings on the wings of
our modern Lepidoptera are always and in all cases unimportant, even
when we cannot interpret their details either as protective, or as a
sign of nauseousness, or as mimetic. The usually very exact similarity
of the colour pattern in the individuals of each species seems to point
to the intervention of personal selection in some form or other, for in
what other way could such a large majority of variations in the same
direction have developed in the germ-plasm as this constancy of the
character indicates.
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