The fact that, as far as our experience goes, superfluous fingers
are never inherited for more than five generations may be simply
explained, for there has been no reason for the intervention of
personal selection, either in the negative sense, for the six-fingered
state does not threaten life, nor in the positive, since it is not of
advantage. The deformity depends on spontaneous germinal variation,
which must have taken place in a majority of ids or it would not
have become manifest. But such a majority of 'polydactylous' ids is
liable to become scattered again in every new descendant, and to be
reduced again into a minority which can no longer make itself felt by
the chances of reducing division and the admixture of normal ids in
amphimixis. A polydactylous race of men could only arise through the
assistance of personal selection; in that case there would doubtless
be just as much chance of success in breeding a six-fingered race as
there was in breeding the crooked-legged Ancon sheep from a single ram
which was malformed in this manner. Without a gradual setting aside of
the germs with normal ids, that is, without personal selection, such
spontaneous deformities, and indeed all _spontaneous_ variations, must
fail of attaining to permanent mastery.
This must frequently be the case in free nature also, but we shall have
to investigate later on, in the section devoted to the formation of
species, whether external circumstances (inbreeding) may not also occur
which make it possible for spontaneous variations to become constant
breed-characters, even although they remain neither good nor bad, and
are thus not subject to the action of personal selection.
In general, however, amphigony with its reduction of the ids and its
constant mingling of strange ids will form the corrective to the
deviations which may arise through the processes of selection within
the id, and which lead to excessive or superfluous development of
certain structures, to a complete disturbance of the harmony of the
parts, and ultimately to the elimination of the species.
Public-domain text, read in full here on John Shaqi.
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