For this reason I am disposed to attribute to saltatory variations a
considerable share in the production of distinctive sexual characters.
From saltatory variations in flowers, fruits, and leaves we know that
these may be conspicuous enough even on their first appearance, and so
we are justified in finding in such variations the first beginnings of
many of the decorative distinguishing characters which occur in the
males of so many animals, especially butterflies and birds. As soon as
it is admitted that variations of considerable amount, which have been
slowly prepared in the germ-plasm by means of germinal selection, can
suddenly attain to expression, one of the objections against sexual
selection is disposed of, for conspicuous variations are necessary for
the operation of this kind of selection, since the changes in question
must attract the attention of the females if they are to be preferred.
Without such preference, even though it be not quite strict and
consistent, a long-continued augmentation of the decorative characters
is inconceivable.
But as intra-germinal disturbances of the position of equilibrium in
the determinant system is at the root of the saltatory variations of
our cultivated plants, it must also have played a large share in the
evolution of breeds among our domesticated animals, which is therefore
by no means wholly due to artificial selection operating upon the
variation of individual characters. In all breeds in the formation of
which the production of more than a single definite character was
concerned, as, for instance, in the broad-nosed breeds of dog--bull-dog
and pug-dog--we may refer the peculiar variation of many parts to
disturbances of the equilibrium of the determinant system, which bring
to light, not suddenly as in the case of saltatory variations, but
gradually and increasingly, the curious complex of characters. Darwin
referred such transformations of the whole animal facies, where a
single varying character is deliberately selected, to correlation, and
by this he understood the mutual influence of the parts of an animal
upon one another. Such correlation certainly exists, as we have already
seen in discussing histonal selection, but here we have rather to do
with the correlation of the parts of the germ-plasm, with the effects
of germinal selection, which, affected by the artificial selection of
particular characters, gradually brings about a more marked disturbance
in the whole determinant system.
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