A few years after the appearance of Fritz Müller's work _Für
Darwin_, Haeckel elaborated Müller's idea, and applied it in a much
more comprehensive manner. He formulated it under the name of 'the
fundamental biogenetic law,' and then he used this 'law' to deduce
from the ontogeny of animals, and more particularly of Man, the
paths of evolution along which our modern species have passed in the
course of the earth's history. In doing so the greatest caution was
necessary, since ontogeny is not an actual unaltered recapitulation
of the phylogeny, but an 'abridged' and in most cases--in my own
belief, in all cases--_a greatly modified recapitulation_. Therefore
we cannot simply accept each ontogenetic stage as an ancestral stage,
but must take into consideration all the facts supplied to us by other
departments of biological inquiry which afford help in the decision
of such questions, especially those brought to light by comparative
morphology and by the whole range of comparative embryology.
Haeckel was quite well aware of this difficulty, and repeatedly
emphasized it by laying stress on the fact that a 'blurring' of the
phyletic stages of development had arisen through the abridgement of
the phylogeny in the ontogeny, and a 'falsification' of it through
the secondary adaptation of individual ontogenetic stages to new
conditions of life. He therefore distinguished between 'Palingenesis,'
that is, simple though abridged repetition of the ancestral history,
and 'Cœnogenesis,' that is, modification of the racial history by
later adaptation of a few or many stages to new conditions of life.
As an example of cœnogenetic modification, I may cite the pupæ of
butterflies. Since these can neither feed nor move from one spot, they
can at no time have been mature forms, and cannot, therefore, represent
independent ancestors of our modern Lepidoptera; they have originated
through the constantly increasing difference between the structure of
the caterpillar and that of the moth or butterfly. Originally, that is,
among the oldest flying insects, the mature animal could be gradually
prepared within the larva as it grew, so that finally nothing was
necessary but a single moult to set free the wings, which had in the
meantime been growing underneath the skin, and to allow the perfect
insect to emerge, complete in all its parts. This is the case even now
with the grasshoppers and crickets. In these forms the larval mode of
life differs very little, if at all, from that of the perfect insect,
and the main difference between the two is the absence of wings in the
larva. But when the perfect insect adapted itself to conditions of life
quite different from the larval conditions, as was the case with the
nectar-sucking bees and butterflies adapted entirely for flight, while
the larvæ were still adapted exclusively to an abundant diet of leaves
and other parts of plants, and to a very inactive life upon plants, the
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