Wiedersheim enumerates nearly a hundred parts thus in process of
degeneration: this means that nearly a hundred structures in Man
are at the present time in process of variation, and this could not
be so unless amphimixis were continually mingling the hereditary
contributions anew from generation to generation, so that the
minus-variations of the parts in question, starting from the germ-plasm
in which they arose at one time as chance variations, and confirmed in
their direction by means of germinal selection, are gradually being
transmitted to all the germ-plasms of the species. We thus see that
even in a period of species-life, which we may fairly call a period of
constancy, variations of a phyletic kind are continually in process,
which could not become general without the co-operation of amphimixis.
Now, we have already seen that personal selection plays no part, or, at
least, no important part in such degenerations, because the variations
which are here concerned do not usually attain to selection value,
but it is just such variations proceeding with infinite slowness that
occur in functionally important organs likewise, and in the progressive
advance of which personal selection and mutual adaptation probably play
a part, so that in this way we can understand why the preservation of
amphigony by natural selection must be effected. It is impossible--for
obvious reasons--to name particular instances with certainty, as we
can do in the case of the rudimentary organs, but even on general
considerations we might expect that among the incipient variations
of the determinants of the germ-plasm there would be some which were
in an ascending direction, and that among these there would be some
which, advanced by germinal selection, would go on ascending until
they attained selection value. Wiedersheim reckons, for instance,
the gradually increasing differentiation of the cortical zone of the
human brain among the parts which are still in process of ascending
variation, and he is probably right in doing so.
But if variations, so slow as to be unnoticeable, are still of abundant
occurrence in Man, we have no reason to doubt that similar processes
are going on in other animals; among the higher Vertebrates at least
there is hardly a species which does not exhibit regressive variations
even now, and in many cases progressive variations also are occurring,
although we cannot give definite proofs of this.
Public-domain text, read in full here on John Shaqi.
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