All these cases are quite in harmony with our conception of amphimixis,
which, now that we have investigated the facts throughout the animate
kingdom, we may sum up in the following propositions. In the whole
organic world, from unicellular organisms up to the highest plants
and animals, amphimixis now means an augmentation of the organism's
power of adaptation to the conditions of its life, since it is only
through amphimixis that simultaneous harmonious adaptation of many
parts becomes possible. It effects this by the mingling and constant
recombination of the germ-plasm ids of different individuals, and thus
gives the selection-processes the chance of favouring advantageous
variational tendencies and eliminating those which are unfavourable,
as well as of collecting and combining all the variations which are
necessary for the further evolution of the species. This indirect
influence of amphimixis on the capacity of organisms for surviving
and being transformed is the fundamental reason for its general
introduction and for its persistence through the whole known realm of
organisms from unicellulars upwards.
The reason for its _first_ introduction among the lower forms of life
must have been a direct effect which had a favourable influence on the
metabolism, and this is so far coincident with the subsequent import of
amphimixis, inasmuch as it may be regarded not only as a heightening of
the power of adaptation, but as an immediate and direct increase and
extension of the power of assimilation. In any case, amphimixis is not
necessary to the actual preservation of life itself, but it does bring
about a wealth and diversity of organic architecture which without it
would have been unattainable.
If amphimixis has been abandoned in the course of phylogeny by isolated
groups of organisms, this has happened because other advantages accrued
to them in consequence, which gave them greater security in the
struggle for existence; but it must be admitted that they thereby lost
their perfect power of adaptation, and that they have thus bartered
their future for the temporary securing of their existence.
In addition to this variational influence, amphimixis has also played a
part in the evolution of sharply defined organic types, especially of
specific types; but of this we shall have more to say later on.
LECTURE XXXI
THE INFLUENCES OF ENVIRONMENT
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