This shows that the cold had affected not only the wing-primordia
of the parental pupæ, but the germ-plasm as well, and at the same
time that this latter variation was less marked than that of the
determinants of the wing-rudiments. This gives rise _to an appearance_
of the transmission of acquired characters.
In the case of many of these cold-aberrations in Lepidoptera the cold
gives rise to variations, but does so not by creating anything new,
but by giving the predominance to primary constituents which have long
been present, but are usually suppressed, and so it is also among the
plants. I have in mind, for instance, the interesting experiments of
Vöchting on the influence of light in the production of flowers in
phanerogams. These showed that the common balsam (_Impatiens noli me
tangere_) produces its familiar open flowers in a strong light, but
in weak light only bears small, closed, so-called 'cleistogamous'
flowers. But it would be utterly erroneous to suppose that the strong
or weak light is the real cause, the _causa materialis_, of these two
forms of flowers: the degree of illumination is merely the stimulus
which provokes one or other of the primary constituents to development,
both kinds being present in the constitution of the plant. As has long
been known, the balsam normally possesses two kinds of flowers, and
the slumbering primary constituents of these are so arranged that the
open flowers develop where there is a prospect of insect visits and
cross-fertilization, that is, in sunny weather or in a strong light,
while closed and inconspicuous flowers adapted for self-fertilization
develop in weak light, that is, in shady places and in concealed parts
of the plant, where insect visits are not to be expected.
[Illustration: FIG. 129. _A_, an aberration of _Arctia caja_, produced
by low temperature. _B_, the most divergent member of its progeny.
After E. Fischer.]
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