Species-colonies of this kind may be due to the agency of man, who has
spread many of his domesticated animals and plants widely over the
earth, but who has also intentionally or unintentionally forced many
wild animals and plants from their original area to distant parts
of the earth, as, for instance, when the English humble-bees were
imported into New Zealand with a view to securing the fertilization
of the clover; but such colonies also occur in thousands of cases
independently of man's agency, and the means by which they are brought
about are very diverse. Little singing-birds are sometimes driven
astray by storms, and carried far away across the sea, to find, if
fortune favours them, a new home on some remote oceanic island;
fresh-water snails, which have just emerged from the egg, creep on to
the broad, webbed feet or among the plumage of a wild duck or some
other migratory bird, and are carried by it far over land and sea,
and finally deposited in a distant marsh or lake. This must happen
not infrequently, as is evidenced by the wide distribution of our
Central European fresh-water snails towards the north and south. But
terrestrial snails can also, though more rarely, be borne in passive
migration far beyond limits which are apparently impassable, as is
evidenced by the presence of land-snails on remote oceanic islands.
The Sandwich Islands are more than 4,000 kilometres from the continent
of America; they originated as volcanoes in the midst of the Pacific
Ocean; and yet they possess a rich fauna of terrestrial snails,
the beginnings of which can only have reached them by the chance
importation of individual snails carried by strayed land-birds.
Charles Darwin was the first who attempted to investigate the problem
of the colonization of oceanic islands by animal inhabitants, and the
chapter in _The Origin of Species_ which deals with the geographical
distribution of animals and plants still forms the basis of all the
investigations directed towards this point. We learn from these that
many land-animals, of which one would not expect it on _a priori_
grounds, may be carried away by chance over the ocean, either, as in
the case of butterflies and other flying insects, and of birds and
bats, by being driven out of their course by the wind, or by being
concealed--either as eggs or as fully-formed animals--in the clefts of
driftwood, where they can resist for a considerable time the usually
destructive influence of salt water. Thus eggs of some of the lowest
Crustacea (Daphnidæ), which are contained in large numbers in the mud
of fresh water, may be transported with some of the mud on the feet
of birds, and this may happen also to encysted infusorians and other
unicellulars, and to the much more highly organized Rotifers as well.
In all these cases, and in many others, it may happen occasionally that
single individuals, or a few at a time, may be carried far afield, and
may reach regions from which their fellows of the same species are
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