The Galapagos Islands lie 1,000 kilometres distant from the coast
of South America, and they too harbour twenty-one endemic species
of land-birds, among them a duck, a buzzard, and about a dozen
different but nearly related mocking-birds, each of which is found
in only one or two of the fifteen islands. The group of islands also
possesses peculiar reptiles, and they take their name from the gigantic
land-tortoises, sometimes 400 kilogrammes in weight, which in Tertiary
times inhabited also the continent of South America, but are now found
in the Galapagos Islands only. The islands also possess endemic lizards
of the genus _Tropidurus_, and although the lizards can no more have
been transported across the ocean than the tortoises, but corroborate
the conclusion drawn from geological data, that the islands were still
connected with the mainland in Tertiary times, the occurrence of a
particular species of _Tropidurus_ upon almost every one of the fifteen
islands testifies anew to the mysterious influence of isolation, for
most of these islands are quite isolated regions for the different
species of lizard, even more than for the mocking-birds, which have
also split up into a series of species.
We are thus led to the hypothesis, which was first introduced into the
Evolution Theory by Darwin, that the prevention of constant crossing
of an isolated colony with the others of the same species from the
original habitat favours the origin of new endemic species, and his
conclusion is confirmed when we learn that islands like the Galapagos
group possess twenty-one endemic land-birds, but only two endemic
sea-birds out of eleven, for the latter traverse great stretches of
sea, and crossings with others of the same species on the neighbouring
continental coasts will often take place. The Bermuda Islands also
afford a proof that the development of endemic species is prevented by
regular crossing with other members of the species from the original
habitat, for although they are 1,200 kilometres distant from the
continent of North America--that is, further than the Galapagos Islands
from South America--they possess no endemic species of bird, and we may
undoubtedly associate this with the fact that the migratory birds from
the continent visit the Bermudas every year.
Madeira also confirms our conclusion, for only one of the ninety-nine
species of bird occurring there can be regarded as endemic, and it has
often been observed that birds from the neighbouring African mainland
(only 240 kilometres distant) are driven across to Madeira. Terrestrial
snails, on the other hand, will seldom be carried to Madeira by birds,
and accordingly we find there an extraordinary number of endemic
terrestrial snails, namely, 109 species.
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