The same thing is shown by the legs and other appendages of insects and
crustaceans, which are adapted for the most diverse functions, and the
individual sections of which must be correlated if the function is to
be possible. Let us consider only the claw structures in crustaceans
and scorpions. Here, too, it seems as if the outgrowth of the last
joint of the leg, which functions as the arm of the claw, must have
arisen as a direct effect of use, through the pressure of an object
held fast by the last joint, the movable half of the claw. Frequently,
moreover, tooth-like protuberances occur on the fixed blade of the
claw (Fig. 103). But how could these have arisen as a direct effect
of pressure, since they are always preformed during the soft state of
the appendage _before use_, and are only made use of after it is fully
hardened. The soft crustaceans, the so-called 'butter-crabs' which
have just cast their shells, creep carefully away and avoid using
their limbs until they have become hard again. Here, too, we have the
co-adaptation of two parts which vary independently, and which cannot
be affected by the Lamarckian principle.
[Illustration: FIG. 103. Claw (_Sch_) on the leg of a 'Beach-fly,' an
Amphipod Crustacean (_Orchestia_). _I_, _II_, the two first joints.
_uA_, the lower blade of the claw, a non-mobile prolongation of the
penultimate joint. _oA_, the upper blade of the claw, the movable
last joint; the tubercles and indentations of the two blades fit one
another. After F. Müller.]
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