represented with excellent effect in another, as not merely a connecting
link in the upward march of progression between the tail of the true fish
and that of the true reptile, but as actually containing in itself—as
the caterpillar contains the future pupa and butterfly—the elements of
the reptilian and mammiferous tail. If there be any virtue in analogy,
the heterocercal tail is, I repeat, of a decidedly higher type than the
homocercal one. It furnishes the first example in the vertebrata of the
coccygeal vertebræ diminishing to a point, which characterizes not only
all the higher reptiles, but also all the higher mammals, and which we
find represented by the _Os coccygis_ in man himself. But to this special
point I shall again refer.
With regard to that rudimentary state of the _occipital_ framework of
the Placoids to which the author of the “Vestiges” refers, it may be but
necessary to say that, notwithstanding the simplicity of their box-like
skulls, they bear in their character, as cases for the protection of
the brain, at least as close an analogy to the skulls of the higher
animals, as those of the osseous fishes, which consist usually of the
extraordinary number of from sixty to eighty bones,—a mark—the author of
the “Vestiges” himself being judge in the case—rather of inferiority than
the reverse. “Elevation is marked in the scale,” we find him saying,
“by an animal exchanging a multiplicity of parts serving one end, for
a smaller number.” The skull of a cod consists of about thrice as many
separate bones as that of a man. But I do not well see that in this case
the fact either of _simplicity_ in excess or of _multiplicity_ in excess
can be insisted upon in either direction, as a proper basis for argument.
Nearly the same remark applies to the maxillaries as to the skull. The
under jaw in man consists of a single bone; that of the thornback—if we
do not include the two suspending _ribs_, which belong equally to the
upper jaw—of two bones, (the number in all the mammiferous quadrupeds:)
that of the cod of four bones, and, if we include the suspending
_ribs_, of twelve. On what principle are we to hold, with _one_ as the
representative number of the highest type of jaw, that _two_ indicates
a lower standing than _four_, or _four_ than _twelve_? In reference to
the further statement, that in many of the ancient fishes “traces can
be observed of the muscles having been attached to the external plates,
strikingly indicating their low grade as vertebrate animals,” it may
be answer enough to state, that the peculiarity in question was not a
characteristic of the _most_ ancient fishes,—the Placoids of the Silurian
system,—but of some Ganoids of the succeeding systems. The reader may
remember, as a case in point, the example furnished by the nail-like bone
of _Asterolepis_, figured in page 111, in which there exists depressions
resembling that of the round ligament in the head of the quadrupedal
thigh-bone.
Public-domain text, read in full here on John Shaqi.
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