displacement of parts_ constitutes yet another form of degradation; and
this form, united, in some instances, to the other two, we find curiously
exemplified in the geological history of the fish,—a history which, with
all its blanks and missing portions, is yet better known than that of
any other division of the vertebrata. And it is, I am convinced, from a
survey of the progress of degradation in the great ichthyic division,—a
progress recorded as “with a pen of iron in the rock for ever,”—and not
from superficial views founded on the cartilaginous or non-cartilaginous
texture of the ichthyic skeleton, that the standing of the kingly fishes
of the earlier periods is to be adequately determined. Any other mode
of survey, save the parallel mode which takes development of brain into
account, evolves, we find, nothing like principle, and lands the inquirer
in inextricable difficulties and inconsistencies.
In all the higher non-degraded vertebrata we find a certain uniform
type of skeleton, consisting of the head, the vertebral column, and
four limbs; and these last, in the various symmetrical forms, whether
exemplified in the higher fish, the higher reptiles, the higher birds,
the higher mammals, or in man himself, occur always in a certain
determinate order. In all the mammals, the scapular bases of the fore
limbs begin opposite the eighth vertebra from the skull backwards, the
seven which go before being cervical or neck vertebræ; in the birds,—a
division of the vertebrata that, from their peculiar organization,
require longer and more flexible necks than the mammals,—the scapulars
begin at distances from the occiput, varying, according to the species,
from opposite the thirteenth to opposite the twenty-fourth vertebra;
and in the reptiles—a division which, according to Cuvier, “presents
a greater diversity of forms, characters, and modes of gait, than any
of the other two,”—they occur at almost all points, from opposite
the second vertebra, as in the frog, to opposite the thirty-third or
thirty-fourth vertebra, as in some species of plesiosaurus. But in
all,—whether mammals, birds, or undegraded reptiles,—they are so placed,
that the creatures possess _necks_, of greater or less length, as an
essential portion of their general type. The hinder limbs have also in
all these three divisions of the animal kingdom their typical place.
They occur opposite, or very nearly opposite, the posterior termination
of the abdominal cavity, and mark the line of separation between the
vertebræ of the trunk (dorsal, lumbar, and sacral) and the third and
last, or _caudal_ division of the column,—a division represented in
man by but four vertebræ, and in the crocodile by about thirty-five,
but which is found to exist, as I have already said, in all the more
perfect forms. The limbs, then, in all the symmetrical animals of the
first three classes of the vertebrata, mark the three great divisions of
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