small depths; whereas the Brachiopoda are deep-sea shells; and the
Cephalopoda may be found voyaging far from land, in the upper strata
of the sea above them. Even in the seas that surround our own island,
the Brachiopodous molluscs—terebratula and crania—have been found, ever
since deep-sea dredging became common, to be not very rare shells; and in
the Mediterranean, where they are less rare still, fleets of Argonauts,
the representatives of a highly organized family of the Cephalopods, to
which it is now believed the Bellerophon of the Palæozoic rocks belonged,
may be seen skimming along the surface, with sail and oar, high over the
profound depths in which they lie. And, of course, when death comes, that
comes to high and low, the remains of both Argonauts and Brachiopods
must lie together at the bottom, in beds almost totally devoid of the
intermediate forms.
Now, the author of the “Vestiges,” in maintaining his hypothesis,
suspends it on the handle furnished him by the immense abundance of the
Silurian Brachiopods. The Silurian period, he says, exhibits “a scanty
and most defective development of life; so much so, that Mr. Lyell calls
it, _par excellence_, the age of Brachiopods, with reference to the by
no means exalted bivalve shell-fish which forms its predominant class.
Such being the actual state of the case, I must persist in describing
even the fauna of this age, which we now know was not the first, as,
generally speaking, such a humble exhibition of the animal kingdom as
we might expect, upon the development theory, to find at an early stage
of the history of organization.” The reader will at once discern the
fallacy here. The Silurian period was peculiarly an age of Brachiopods,
for in no other period were Brachiopods so numerous, specifically or
individually, or of such size or importance; whereas it was not _so
peculiarly_ an age of Cephalopods, for these we find introduced in still
greater numbers during the Liasic and Oolitic periods. In 1848, when
Professor Edward Forbes edited the Palæontological map of Britain and
Ireland, which forms one of the very admirable series of “Johnstone’s
Physical Atlas,” the Cephalopods of the Silurian rocks of England and
Wales were estimated at forty-eight species, and the Brachiopods at
one hundred and fifty; whereas at the same date there were two hundred
and five Cephalopods of the Oolitic formations enumerated, and but
fifty-four Brachiopods. It is the molluscs of the inferior, not those
of the superior class, that constitute (with their contemporaries the
Trilobites) the characteristic fossils of the Silurian rocks; and hence
the propriety of the distinctive name suggested by Sir Charles Lyell.
But in the development question, what we have specially to consider is,
not the _numbers_ of the low, but the _standing_ of the high. A country
may be distinctively a country of flocks and herds, or a country of
the carnivorous mammalia, or, like New South Wales or the Galapagos, a
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