more of each scale covered than exposed; and this is so rare a ganoidal
mode of arrangement, that, with the exception of the _Dipterus_,—a genus
which, though it gives its name to the Dipterian sept, differed greatly
from every other Dipterian,—I know not, beyond the limits of the ancient
Cœlacanth family, a single Ganoid that possessed it. The bony covering of
the Cœlacanths was _farthest_ removed in character from shagreen, as that
of their contemporaries the Acanths approximated to it most nearly; they
were, in this respect, the two extremes of their order; and did we find
the Cœlacanths in but the later geological formations, while the Acanths
were restricted to the earlier, it might be argued by assertors of the
development hypothesis, that the amply imbricated, slate-like scale of
the latter had been developed in the lapse of ages from the shagreen
tubercle, by passing in its downward course—broadening and expanding
as it descended—through the minute, scarcely imbricated disks of the
Acanths, and the more amply imbricated tile-like rhombs of the Dipterians
and Palæonisci, until it had reached its full extent of imbrication in
the familiar modern type exemplified in both the Cœlacanths and the
ordinary fishes. But such is not the order which nature has observed;—the
two extremes of the ganoid scale appear together in the same early
formation: both become extinct at a period geologically remote; and the
ganoid scales of the existing state of things which most nearly resemble
those of ancient time are scales formed on the intermediate or tile-roof
principle.
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