The cranial buckler of the _Dipterus_ (fig. 19) was, like that of the
_Diplopterus_, of great beauty. In some of the finer specimens, we find
the enamel ornately tatooed, within the more strongly-marked divisions,
by delicately traced lines, waved and bent, as if upon the principle
of Hogarth; and though the lateral plates are numerous and small, and
defy the homologies, we may trace in those of the central line, from the
snout to the nape, what seem to be the representatives of the frontal,
parietal, and occipital bones,—the parietals ranging, as in the skull
of the carp and in that of most of the mammals, in their proper place
in the medial line. But the under surface of the cranium, armed, as on
the upper surface, with plates of bone, exhibited an arrangement still
more peculiar, (fig. 20.) Its rectangular patches of palatal teeth, its
curious dart-like bone, placed immediately behind these, and attached,
as the dart-head is attached to the handle, to a broad lozenge-shaped
plate, with two strong osseous processes projecting on either side, forms
such a _tout ensemble_ as is unique among fishes. Even here, however,
there may be traced at least a shade of homological resemblance to the
bones which form the base of the osseous skull. The single lozenge-shaped
plate, (A,) with its dart-head, occupies the place of the basi-occipital
bone; the posterior portion of the vomer seems represented by a strong
bony ridge, extending towards the snout; two separate bones, each bearing
one of the angular patches of teeth, corresponds to the sphenoid bone
and its alæ; and attached laterally to each of these there is the strong
projecting bone, on which the lower jaw appears to have hinged, and
which apparently represents the lower part of the temporal bone. Not less
singular was the form of the creature’s under jaw, (fig. 21.) I know no
other fish-jaw, whether of the recent or the extinct races, that might
be so readily mistaken for that of a quadruped. It exhibits not only the
condyloid, but also the coronoid processes; and, save that it broadens
on its upper edges, where in mammals the grinders are placed, so as to
furnish field enough for angular patches of teeth, which correspond with
the angular patches in the palate, it might be regarded, found detached,
as at least a reptilian, if not mammalian, bone. The disposition of
the palatal teeth of the _Dipterus_ will scarce fail to remind the
mechanist of the style of grooving resorted to in the formation of
mill-stones for the grinding of flour; nor is it wholly improbable that,
in correspondence with the rotatory motion of the stones to which the
grooving is specially adapted, jaws so hinged may have possessed some
such power of lateral motion as that exemplified by the human subject in
the use of the molar teeth.
[Illustration: Fig. 21.
UNDER JAW OF DIPTERUS.]
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