These cranial bucklers, extending from nape to snout, protected the
exposed upper surface of the cartilaginous skull, and conformed to it in
shape, as a helmet conforms to the shape of the head, or a breast-plate
to the shape of the chest. And as the cartilaginous heads resembled in
general outline the osseous ones, the buckler which covered their upper
surface resembled in general outline the upper surface of the osseous
skull. It was in no case entirely a flat plate; but in every species
rounded over the snout and in most species at the sides; and so, in
order that its characteristic proportions might be preserved throughout
the various stages of growth in the head which it covered, it had to be
formed from several distinct centres of ossification, and to extend in
area around the edges of the plates originated from these. The workman
finds no difficulty in adding to the size of a piece of straight wall,
whether by heightening or lengthening it; but he cannot add to the size
of a dome or arch, without first taking it down, and then erecting it
anew on a larger scale. In the domes and arches of the animal kingdom,
the problem is solved by building them up of distinct pieces, few or
many, according to the demands of the figure which they compose, and
rendering these pieces capable of increase along their edges. It is on
this principle that the Cystidea, the Echinidæ, the Chelonian carapace
and plastron, and the skulls of the osseous Vertebrata, are constructed.
It is also the principle on which the cranial bucklers of the ancient
Ganoids were formed.[17] And from the general resemblance in figure of
these bucklers to the upper surface of the osseous skull, the separate
parts necessary for the building up of the one were anticipated, by many
ages, in the building up of the other; just as we find external arches
of stone which were erected two thousand years ago, constructed on the
same principle, and relatively of the same parts, as internal arches of
brick built in the present age. Doubtless, however, with this mechanical
necessity for correspondence of parts in the formation of corresponding
erections, there may have mingled that regard for typical resemblance
which seems so marked a characteristic of the _style_, if I may so
express myself, in which the Divine Architect gives expression to his
ideas. The external osseous buckler He divided after the general pattern
which was to be exemplified, in latter times, in the divisions of the
internal osseous skull; as if in illustration of that “ideal exemplar”
which dwelt in his mind from eternity, and on the palpable existence
of which sober science has based deductions identical in their scope
and bearing with some of the sublimest doctrines of the theologian.
“The recognition,” says Professor Owen, “of an ideal exemplar for the
vertebrated animals, proves that the knowledge of such a being as man
existed before man appeared; for the Divine mind which planned the
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