to its apparent homologue, the radius. Fig. _e_, a slim light bone,
which narrows and thickens in the centre, and flattens and broadens at
each end, was probably a scapula or shoulder-blade,—a bone which in
most fishes _splices_ on, as a sailor would say, by squamose jointings,
to the coracoidian bone at the one end, and the super-scapular bone at
the other. As indicated by its size, it must have belonged to a small
individual: it is, however, twice as long, and about six times as bulky,
as the scapula of a large cod.
[Illustration: Fig. 41.
INTERNAL BONES OF ASTEROLEPIS.
(One third nat. size, linear.)]
[Illustration: Fig. 42.
ISCHIUM OF ASTEROLEPIS.
(One half nat. size, linear.)]
Of the bone represented in fig. 42, I have determined, from a Cromarty
specimen, the place and use: it formed the interior base to which
one of the ventral fins was attached. In all fishes the bones of the
hinder extremities are inadequately represented: in none do we find the
pelvic arch complete; and to that nether portion of it which we do find
represented, and which Professor Owen regards as the homologue of the
_os ischium_ or hip-bone, the homologues of the metatarsal and toe-bones
are attached, to the exclusion of the bones of the thigh and leg. In the
Abdominales,—fishes such as the salmon and carp,—that have the ventrals
placed behind the abdomen, in the position analogous to that in which
the hinder legs of the reptiles and mammals occur, the ischiatic bones
generally exist as flat triangular plates, with their heads either
turned _inwards_ and downwards, as in the herring, or _outwards_ and
downwards, as in the pike; whereas in some of the cartilaginous fishes,
such as the Rays and Sharks, they exist as an undivided cartilaginous
band, stretched transversely from ventral to ventral. And such, with
but an upward direction, appears to have been their position in the
_Asterolepis_. They seem to have united at the narrow neck A, over the
middle of the lower portion of the abdomen; and to the notches of the
flat expansion B,—notches which exactly resemble those of the immensely
developed carpal bones of the Ray,—five metatarsal bones were attached,
from which the fin expanded. It is interesting to find the number in this
ancient representative of the vertebrata restricted to five,—a number
greatly exceeded in most of the existing fishes, but which is the true
normal number of the vertebrate sub-kingdom as shown in all the higher
examples such as man, the _quadrumana_, and in most of the _carnaria_.
The form of this bone somewhat resembles that of the analogous bone in
those fishes, such as the perch and gurnard, cod and haddock, which have
their ventrals suspended to the scapular belt; but its position in the
Cromarty specimen, and that of the ventrals in the various specimens of
the Cœlacanth family in which their place is still shown, forbids the
supposition that _it_ was so suspended,—a circumstance in keeping with
Public-domain text, read in full here on John Shaqi.
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