plate can be inscribed—such is its form—in a semicircle, of which the
nail-shaped ridge in the middle (if we strike off a minute portion of the
sharp point, usually wanting in detached specimens) forms very nearly
the radius, and of which the diameter equals the breadth of the cranial
buckler, along a line drawn across at a distance from the nape, equal
to two thirds of the distance between the nape and the eyes. Thus, the
largest diameter of a hyoid plate which belonged to a cranial buckler a
foot in length is, I find, equal to seven one quarter inches, while the
length of its nape somewhat exceeds three five eighth inches. The nail of
the Stromness specimen measures five and a half inches. It must have run
along a hyoid plate eleven inches in transverse breadth, and have been
associated with a cranial buckler eighteen one eighth inches in length;
and the _Asterolepis_ to which it belonged must have measured from snout
to tail, if formed, as it probably was, in the proportions of its brother
Cœlacanth the _Glyptolepis_, eight feet three inches; and if in those
of the _Diplopterus_, from nine feet nine to ten feet six inches. This
oldest of Scottish fish—this earliest-born of the Ganoids yet known—was
at least as bulky as a large porpoise.
It was small, however, compared with specimens of the _Asterolepis_
found elsewhere. The hyoid plate figured in page 110, (fig. 36,)—a
Thurso specimen which I owe to the kindness of Mr. Dick,—measures
nearly fourteen inches, and the cranial buckler of the same individual,
fifteen one fourth inches, in breadth. The latter, when entire, must
have measured twenty-three one half inches in length; and the fish to
which it belonged, if formed in the proportions of the _Glyptolepis_,
ten feet six inches; and if in those of the _Diplopterus_, from twelve
feet five to thirteen feet eight inches in length. Did the shield still
exist in its original state as a buckler of tough, enamel-crusted bone,
it might be converted into a Highland target, nearly broad enough to
cover the ample chest of a Rob Roy or Allan M’Aulay, and strong enough to
dash aside the keenest broadsword. Another hyoid plate found by Mr. Dick
measures sixteen one half inches in breadth; and a cast in the British
Museum, from one of the Russian specimens of Professor Asmus, (fig. 46,)
twenty-four inches. The individual to which this last plate belonged
must, if built in the shorter proportions, have measured eighteen, and if
in the longer, twenty-three feet in length. The two hyoid plates of the
specimen of _Holoptychius_ in the British Museum measure but four and a
half inches along that transverse line in which the Russian _Asterolepis_
measures two feet, and the largest Thurso specimen sixteen inches and a
half. The maxillary bone of a cod-fish two and a half feet from snout to
tail measures three inches in length. One of the Russian maxillary bones
in the possession of Professor Asmus measures in length twenty-eight
inches.
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