The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
and where it was less than 1° some evidence of their flowing down could
still be detected. On another occasion, after rain which was never
heavy, but which lasted for 18 hours, all the castings on this same
gently inclined lawn had lost their vermiform structure; and they had
flowed, so that fully two-thirds of the ejected earth lay below the
mouths of the burrows.
These observations led me to make others with more care. Eight castings
were found on my lawn, where the grass-blades are fine and close
together, and three others on a field with coarse grass. The
inclination of the surface at the eleven places where these castings
were collected varied between 4° 30′ and 17° 30′; the mean of the
eleven inclinations being 9° 26′. The length of the castings in the
direction of the slope was first measured with as much accuracy as
their irregularities would permit. It was found possible to make these
measurements within about ⅛ of an inch, but one of the castings was too
irregular to admit of measurement. The average length in the direction
of the slope of the remaining ten castings was 2.03 inches. The
castings were then divided with a knife into two parts along a
horizontal line passing through the mouth of the burrow, which was
discovered by slicing off the turf; and all the ejected earth was
separately collected, namely, the part above the hole and the part
below. Afterwards these two parts were weighed. In every case there was
much more earth below than above; the mean weight of that above being
103 grains, and of that below 205 grains; so that the latter was very
nearly double the former. As on level ground castings are commonly
thrown up almost equally round the mouths of the burrows, this
difference in weight indicates the amount of ejected earth which had
flowed down the slope. But very many more observations would be
requisite to arrive at any general result; for the nature of the
vegetation and other accidental circumstances, such as the heaviness of
the rain, the direction and force of the wind, &c., appear to be more
important in determining the quantity of the earth which flows down a
slope than its angle. Thus with four castings on my lawn (included in
the above eleven) where the mean slope was 7° 19′, the difference in
the amount of earth above and below the burrows was greater than with
three other castings on the same lawn where the mean slope was 12° 5′.
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