The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
analyses, only from 1 to 2 per cent. of earthy matter; and two samples
from pits near my house contained 1.3 and 0.6 per cent. I mention these
latter cases because, from the thickness of the overlying bed of red clay
with flints, I had imagined that the underlying chalk might here be less
pure than elsewhere. The cause of the residue accumulating more in some
places than in others, may be attributed to a layer of argillaceous
matter having been left at an early period on the chalk, and this would
check the subsequent percolation of earthy matter into it.
From the facts now given we may conclude that castings ejected on our
Chalk Downs suffer some loss by the percolation of their finer matter
into the chalk. But such impure superficial chalk, when dissolved, would
leave a larger supply of earthy matter to be added to the mould than in
the case of pure chalk. Besides the loss caused by percolation, some
fine earth is certainly washed down the sloping grass-covered surfaces of
our Downs. The washing-down process, however, will be checked in the
course of time; for although I do not know how thin a layer of mould
suffices to support worms, yet a limit must at last be reached; and then
their castings would cease to be ejected or would become scanty.
The following cases show that a considerable quantity of fine earth is
washed down. The thickness of the mould was measured at points 12 yards
apart across a small valley in the Chalk near Winchester. The sides
sloped gently at first; then became inclined at about 20°; then more
gently to near the bottom, which transversely was almost level and about
50 yards across. In the bottom, the mean thickness of the mould from
five measurements was 8.3 inches; whilst on the sides of the valley,
where the inclination varied between 14° and 20°, its mean thickness was
rather less than 3.5 inches. As the turf-covered bottom of the valley
sloped at an angle of only between 2° and 3°, it is probable that most of
the 8.3-inch layer of mould had been washed down from the flanks of the
valley, and not from the upper part. But as a shepherd said that he had
seen water flowing in this valley after the sudden thawing of snow, it is
possible that some earth may have been brought down from the upper part;
or, on the other hand, that some may have been carried further down the
valley. Closely similar results, with respect to the thickness of the
mould, were obtained in a neighbouring valley.
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