The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
Worms are poorly provided with sense-organs, for they cannot be said to
see, although they can just distinguish between light and darkness; they
are completely deaf, and have only a feeble power of smell; the sense of
touch alone is well developed. They can therefore learn but little about
the outside world, and it is surprising that they should exhibit some
skill in lining their burrows with their castings and with leaves, and in
the case of some species in piling up their castings into tower-like
constructions. But it is far more surprising that they should apparently
exhibit some degrees of intelligence instead of a mere blind instinctive
impulse, in their manner of plugging up the mouths of their burrows.
They act in nearly the same manner as would a man, who had to close a
cylindrical tube with different kinds of leaves, petioles, triangles of
paper, &c., for they commonly seize such objects by their pointed ends.
But with thin objects a certain number are drawn in by their broader
ends. They do not act in the same unvarying manner in all cases, as do
most of the lower animals; for instance, they do not drag in leaves by
their foot-stalks, unless the basal part of the blade is as narrow as the
apex, or narrower than it.
* * * * *
When we behold a wide, turf-covered expanse, we should remember that its
smoothness, on which so much of its beauty depends, is mainly due to all
the inequalities having been slowly levelled by worms. It is a
marvellous reflection that the whole of the superficial mould over any
such expanse has passed, and will again pass, every few years through the
bodies of worms. The plough is one of the most ancient and most valuable
of man’s inventions; but long before he existed the land was in fact
regularly ploughed, and still continues to be thus ploughed by
earth-worms. It may be doubted whether there are many other animals
which have played so important a part in the history of the world, as
have these lowly organized creatures. Some other animals, however, still
more lowly organized, namely corals, have done far more conspicuous work
in having constructed innumerable reefs and islands in the great oceans;
but these are almost confined to the tropical zones.
FOOTNOTES.
{2} ‘Leçons de Géologie Pratique,’ tom. i. 1845, p. 140.
{3} ‘Transactions Geolog. Soc.’ vol. v. p. 505. Read November 1, 1837.
{4a} ‘Histoire des progrès de la Géologie,’ tom. i. 1847, p. 224.
{4b} ‘Zeitschrift für wissenschaft. Zoologie,’ B. xxviii. 1877, p. 361.
{5} ‘Gardeners’ Chronicle,’ April 17, 1869, p. 418.
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