The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
WEIGHT OF THE CASTINGS ACCUMULATED AT THE MOUTH OF A SINGLE BURROW.
(1.) Down, Kent (sub-soil red clay, full of flints, 3.98
over-lying the chalk). The largest casting which I could
find on the flanks of a steep valley, the sub-soil being
here shallow. In this one case, the casting was not well
dried
(2.) Down.—Largest casting which I could find (consisting 3.87
chiefly of calcareous matter), on extremely poor pasture
land at the bottom of the valley mentioned under (1.)
(3.) Down.—A large casting, but not of unusual size, from 1.22
a nearly level field, poor pasture, laid down in a grass
about 35 years before
(4.) Down. Average weight of 11 not large castings 0.7
ejected on a sloping surface on my lawn, after they had
suffered some loss of weight from being exposed during a
considerable length of time to rain
(5.) Near Nice in France.—Average weight of 12 castings of 1.37
ordinary dimensions, collected by Dr. King on land which
had not been mown for a long time and where worms abounded,
viz., a lawn protected by shrubberies near the sea; soil
sandy and calcareous; these castings had been exposed for
some time to rain, before being collected, and must have
lost some weight by disintegration, but they still retained
their form
(6.) The heaviest of the above twelve castings 1.76
(7.) Lower Bengal.—Average weight of 22 castings, 1.24
collected by Mr. J. Scott, and stated by him to have been
thrown up in the course of one or two nights
(8.) The heaviest of the above 22 castings 2.09
(9.) Nilgiri Mountains, S. India; average weight of the 5 3.15
largest castings collected by Dr. King. They had been
exposed to the rain of the last monsoon, and must have lost
some weight
(10.) The heaviest of the above 5 castings 4.34
In this table we see that castings which had been ejected at the mouth of
the same burrow, and which in most cases appeared fresh and always
retained their vermiform configuration, generally exceeded an ounce in
weight after being dried, and sometimes nearly equalled a quarter of a
pound. On the Nilgiri mountains one casting even exceeded this latter
weight. The largest castings in England were found on extremely poor
pasture-land; and these, as far as I have seen, are generally larger than
those on land producing a rich vegetation. It would appear that worms
have to swallow a greater amount of earth on poor than on rich land, in
order to obtain sufficient nutriment.
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