The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
The second square was marked on unenclosed common land, at a height of
about 700 ft. above the sea, at some little distance from Leith Hill
Tower. The surface was clothed with short, fine turf, and had never been
disturbed by the hand of man. The spot selected appeared neither
particularly favourable nor the reverse for worms; but I have often
noticed that castings are especially abundant on common land, and this
may, perhaps, be attributed to the poorness of the soil. The vegetable
mould was here between three and four inches in thickness. As this spot
was at some distance from the house where the lady lived, the castings
were not collected at such short intervals of time as those on the
terrace; consequently the loss of fine earth during rainy weather must
have been greater in this than in the last case. The castings moreover
were more sandy, and in collecting them during dry weather they sometimes
crumbled into dust, and much was thus lost. Therefore it is certain that
the worms brought up to the surface considerably more earth than that
which was collected. The last collection was made on October 27th, 1871;
i.e., 367 days after the square had been marked out and the surface
cleared of all pre-existing castings. The collected castings, after
being well dried, weighed 7.453 pounds; and this would give, for an acre
of the same kind of land, 16.1 tons of annually ejected dry earth.
SUMMARY OF THE FOUR FOREGOING CASES.
(1.) Castings ejected near Nice within about a year, collected by Dr.
King on a square foot of surface, calculated to yield per acre 14.58
tons.
(2.) Castings ejected during about 40 days on a square yard, in a
field of poor pasture at the bottom of a large valley in the Chalk,
calculated to yield annually per acre 18.12 tons.
(3.) Castings collected from a square yard on an old terrace at Leith
Hill Place, during 369 days, calculated to yield annually per acre
7.56 tons.
(4.) Castings collected from a square yard on Leith Hill Common
during 367 days, calculated to yield annually per acre 16.1 tons.
_The thickness of the layer of mould_, _which castings ejected during a
year would form if uniformly spread out_.—As we know, from the two last
cases in the above summary, the weight of the dried castings ejected by
worms during a year on a square yard of surface, I wished to learn how
thick a layer of ordinary mould this amount would form if spread
uniformly over a square yard. The dry castings were therefore broken
into small particles, and whilst being placed in a measure were well
shaken and pressed down. Those collected on the Terrace amounted to
124.77 cubic inches; and this amount, if spread out over a square yard,
would make a layer 0.9627 inch in thickness. Those collected on the
Common amounted to 197.56 cubic inches, and would make a similar layer
0.1524 inch in thickness.
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