The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
In these cases (excepting the last) it may be seen that the amount of
earth brought to the surface during 10 years is somewhat greater than
that calculated from the castings which were actually weighed. This
excess may be partly accounted for by the loss which the weighed castings
had previously undergone through being washed by rain, by the adhesion of
particles to the blades of the surrounding grass, and by their crumbling
when dry. Nor must we overlook other agencies which in all ordinary
cases add to the amount of mould, and which would not be included in the
castings that were collected, namely, the fine earth brought up to the
surface by burrowing larvæ and insects, especially by ants. The earth
brought up by moles generally has a somewhat different appearance from
vegetable mould; but after a time would not be distinguishable from it.
In dry countries, moreover, the wind plays an important part in carrying
dust from one place to another, and even in England it must add to the
mould on fields near great roads. But in our country these latter
several agencies appear to be of quite subordinate importance in
comparison with the action of worms.
We have no means of judging how great a weight of earth a single
full-sized worm ejects during a year. Hensen estimates that 53,767 worms
exist in an acre of land; but this is founded on the number found in
gardens, and he believes that only about half as many live in
corn-fields. How many live in old pasture land is unknown; but if we
assume that half the above number, or 26,886 worms live on such land,
then taking from the previous summary 15 tons as the weight of the
castings annually thrown up on an acre of land, each worm must annually
eject 20 ounces. A full-sized casting at the mouth of a single burrow
often exceeds, as we have seen, an ounce in weight; and it is probable
that worms eject more than 20 full-sized castings during a year. If they
eject annually more than 20 ounces, we may infer that the worms which
live in an acre of pasture land must be less than 26,886 in number.
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