The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their HabitsDarwin, Charles
Science
The Formation of Vegetable Mould Through the Action of Worms: With Observations on Their Habits
Darwin, Charles
Earthworms; Humification; Soils
Aug. 26th, 1877; that is, five days after the floor had been cleared. On
the previous night there had been some heavy rain, which washed the
surface clean, and now the mouths of forty burrows were counted. Parts
of the concrete were seen to be solid, and had never been penetrated by
worms, and here the rain-water lodged.
Sept. 5th.—Tracks of worms, made during the previous night, could be seen
on the surface of the floor, and five or six vermiform castings had been
thrown up. These were defaced.
Sept. 12th.—During the last six days, the worms have not been active,
though many castings have been ejected in the neighbouring fields; but on
this day the earth was a little raised over the mouths of the burrows, or
castings were ejected, at ten fresh points. These were defaced. It
should be understood that when a fresh burrow is spoken of, this
generally means only that an old burrow has been re-opened. Mr. Farrer
was repeatedly struck with the pertinacity with which the worms re-opened
their old burrows, even when no earth was ejected from them. I have
often observed the same fact, and generally the mouths of the burrows are
protected by an accumulation of pebbles, sticks or leaves. Mr. Farrer
likewise observed that the worms living beneath the floor of the atrium
often collected coarse grains of sand, and such little stones as they
could find, round the mouths of their burrows.
Sept. 13th; soft wet weather. The mouths of the burrows were re-opened,
or castings were ejected, at 31 points; these were all defaced.
Sept. 14th; 34 fresh holes or castings; all defaced.
Sept. 15th; 44 fresh holes, only 5 castings; all defaced.
Sept. 18th; 43 fresh holes, 8 castings; all defaced.
The number of castings on the surrounding fields was now very large.
Sept. 19th; 40 holes, 8 castings; all defaced.
Sept. 22nd; 43 holes, only a few fresh castings; all defaced.
Sept. 23rd; 44 holes, 8 castings.
Sept. 25th; 50 holes, no record of the number of castings.
Oct. 13th; 61 holes, no record of the number of castings.
After an interval of three years, Mr. Farrer, at my request, again looked
at the concrete floor, and found the worms still at work.
Knowing what great muscular power worms possess, and seeing how soft the
concrete was in many parts, I was not surprised at its having been
penetrated by their burrows; but it is a more surprising fact that the
mortar between the rough stones of the thick walls, surrounding the
rooms, was found by Mr. Farrer to have been penetrated by worms. On
August 26th, that is, five days after the ruins had been exposed, he
observed four open burrows on the broken summit of the eastern wall (W in
Fig. 8); and, on September 15th, other burrows similarly situated were
seen. It should also be noted that in the perpendicular side of the
trench (which was much deeper than is represented in Fig. 8) three recent
burrows were seen, which ran obliquely far down beneath the base of the
old wall.
Public-domain text, read in full here on John Shaqi.
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