{311} _Origin_, Ed. i. p. 301, vi. p. 440. {312} _Origin_, Ed. i. p. 329, vi. p. 471. {313} The structure of the Pachyderm leg was a favourite with the author. It is discussed in the Essay of 1842, p. 48. In the present Essay the following sentence in the margin appears to refer to Pachyderms and Ruminants: "There can be no doubt, if we banish all fossils, existing groups stand more separate." The following occurs between the lines "The earliest forms would be such as others could radiate from." As I have admitted the high improbability, that if every fossil were disinterred, they would compose in each of the Divisions of Nature a perfect series of the kind required; consequently I freely admit, that if those geologists are in the right who consider the lowest known formation as contemporaneous with the first appearances of life{314}; or the several formations as at all closely consecutive; or any one formation as containing a nearly perfect record of the organisms which existed during the whole period of its deposition in that quarter of the globe;--if such propositions are to be accepted, my theory must be abandoned. {314} _Origin_, Ed. i. p. 307, vi. p. 448. If the Palaeozoic system is really contemporaneous with the first appearance of life, my theory must be abandoned, both inasmuch as it limits _from shortness of time_ the total number of forms which can have existed on this world, and because the organisms, as fish, mollusca{315} and star-fish found in its lower beds, cannot be considered as the parent forms of all the successive species in these classes. But no one has yet overturned the arguments of Hutton and Lyell, that the lowest formations known to us are only those which have escaped being metamorphosed <illegible>; if we argued from some considerable districts, we might have supposed that even the Cretaceous system was that in which life first appeared. From the number of distant points, however, in which the Silurian system has been found to be the lowest, and not always metamorphosed, there are some objections to Hutton's and Lyell's view; but we must not forget that the now existing land forms only 1/5 part of the superficies of the globe, and that this fraction is only imperfectly known. With respect to the fewness of the organisms found in the Silurian and other Palaeozoic formations, there is less difficulty, inasmuch as (besides their gradual obliteration) we can expect formations of this vast antiquity to escape entire denudation, only when they have been accumulated over a wide area, and have been subsequently protected by vast superimposed deposits: now this could generally only hold good with deposits accumulating in a wide and deep ocean, and therefore unfavourable to the presence of many living things. A mere narrow and not very thick strip of matter, deposited along a coast where organisms most abound, would have no chance of escaping denudation and