The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
In the Tertiary system the same facts, which make us admit as probable
that new species have slowly appeared, lead to the admission that old
ones have slowly disappeared, not several together, but one after
another; and by analogy one is induced to extend this belief to the
Secondary and Palaeozoic epochs. In some cases, as the subsidence of a
flat country, or the breaking or the joining of an isthmus, and the
sudden inroad of many new and destructive species, extinction might be
locally sudden. The view entertained by many geologists, that each fauna
of each Secondary epoch has been suddenly destroyed over the whole
world, so that no succession could be left for the production of new
forms, is subversive of my theory, but I see no grounds whatever to
admit such a view. On the contrary, the law, which has been made out,
with reference to distinct epochs, by independent observers, namely,
that the wider the geographical range of a species the longer is its
duration in time, seems entirely opposed to any universal
extermination{330}. The fact of species of mammiferous animals and fish
being renewed at a quicker rate than mollusca, though both aquatic; and
of these the terrestrial genera being renewed quicker than the marine;
and the marine mollusca being again renewed quicker than the Infusorial
animalcula, all seem to show that the extinction and renewal of species
does not depend on general catastrophes, but on the particular relations
of the several classes to the conditions to which they are exposed{331}.
{330} Opposite to this passage the author has written "d'Archiac,
Forbes, Lyell."
{331} This passage, for which the author gives as authorities the
names of Lyell, Forbes and Ehrenberg, corresponds in part to the
discussion beginning on p. 313 of _Origin_, Ed. i., vi. p. 454.
Some authors seem to consider the fact of a few species having
survived{332} amidst a number of extinct forms (as is the case with a
tortoise and a crocodile out of the vast number of extinct sub-Himalayan
fossils) as strongly opposed to the view of species being mutable. No
doubt this would be the case, if it were presupposed with Lamarck that
there was some inherent tendency to change and development in all
species, for which supposition I see no evidence. As we see some species
at present adapted to a wide range of conditions, so we may suppose that
such species would survive unchanged and unexterminated for a long time;
time generally being from geological causes a correlative of changing
conditions. How at present one species becomes adapted to a wide range,
and another species to a restricted range of conditions, is of difficult
explanation.
{332} The author gives Falconer as his authority: see _Origin_, Ed.
i. p. 313, vi. p. 454.
_Extinction of species._
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