The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
We will now turn to the floras of mountain-summits which are well known
to differ from the floras of the neighbouring lowlands. In certain
characters, such as dwarfness of stature, hairiness, &c., the species
from the most distant mountains frequently resemble each other,--a kind
of analogy like that for instance of the succulency of most desert
plants. Besides this analogy, Alpine plants present some eminently
curious facts in their distribution. In some cases the summits of
mountains, although immensely distant from each other, are clothed by
the same identical species{362} which are likewise the same with those
growing on the likewise very distant Arctic shores. In other cases,
although few or none of the species may be actually identical, they are
closely related; whilst the plants of the lowland districts surrounding
the two mountains in question will be wholly dissimilar. As
mountain-summits, as far as their plants are concerned, are islands
rising out of an ocean of land in which the Alpine species cannot live,
nor across which is there any known means of transport, this fact
appears directly opposed to the conclusion which we have come to from
considering the general distribution of organisms both on continents and
on islands--namely, that the degree of relationship between the
inhabitants of two points depends on the completeness and nature of the
barriers between those points{363}. I believe, however, this anomalous
case admits, as we shall presently see, of some explanation. We might
have expected that the flora of a mountain summit would have presented
the same relation to the flora of the surrounding lowland country, which
any isolated part of a continent does to the whole, or an island does to
the mainland, from which it is separated by a rather wide space of sea.
This in fact is the case with the plants clothing the summits of _some_
mountains, which mountains it may be observed are particularly isolated;
for instance, all the species are peculiar, but they belong to the forms
characteristic of the surrounding continent, on the mountains of
Caraccas, of Van Dieman's Land and of the Cape of Good Hope{364}. On
some other mountains, for instance <in> Tierra del Fuego and in Brazil,
some of the plants though distinct species are S. American forms; whilst
others are allied to or are identical with the Alpine species of Europe.
In islands of which the lowland flora is distinct <from> but allied to
that of the nearest continent, the Alpine plants are sometimes (or
perhaps mostly) eminently peculiar and distinct{365}; this is the case
on Teneriffe, and in a lesser degree even on some of the Mediterranean
islands.
{362} See _Origin_, Ed. i. p. 365, vi. p. 515. The present
discussion was written before the publication of Forbes' celebrated
paper on the same subject; see _Life and Letters_, vol. I. p. 88.
Public-domain text, read in full here on John Shaqi.
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