The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
The manner in which, according to our theory, the arrangement of species
in groups is due to partial extinction, will perhaps be rendered clearer
in the following way. Let us suppose in any one great class, for
instance in the Mammalia, that every species and every variety, during
each successive age, had sent down one unaltered descendant (either
fossil or living) to the present time; we should then have had one
enormous series, including by small gradations every known mammiferous
form; and consequently the existence of groups{450}, or chasms in the
series, which in some parts are in greater width, and in some of less,
is solely due to former species, and whole groups of species, not having
thus sent down descendants to the present time.
{450} The author probably intended to write "groups separated by
chasms."
With respect to the "analogical" or "adaptive" resemblances between
organic beings which are not really related{451}, I will only add, that
probably the isolation of different groups of species is an important
element in the production of such characters: thus we can easily see, in
a large increasing island, or even a continent like Australia, stocked
with only certain orders of the main classes, that the conditions would
be highly favourable for species from these orders to become adapted to
play parts in the economy of nature, which in other countries were
performed by tribes especially adapted to such parts. We can understand
how it might happen that an otter-like animal might have been formed in
Australia by slow selection from the more carnivorous Marsupial types;
thus we can understand that curious case in the southern hemisphere,
where there are no auks (but many petrels), of a petrel{452} having been
modified into the external general form so as to play the same office
in nature with the auks of the northern hemisphere; although the habits
and form of the petrels and auks are normally so wholly different. It
follows, from our theory, that two orders must have descended from one
common stock at an immensely remote epoch; and we can perceive when a
species in either order, or in both, shows some affinity to the other
order, why the affinity is usually generic and not particular--that is
why the Bizcacha amongst Rodents, in the points in which it is related
to the Marsupial, is related to the whole group{453}, and not
particularly to the Phascolomys, which of all Marsupialia is related
most to the Rodents. For the Bizcacha is related to the present
Marsupialia, only from being related to their common parent-stock; and
not to any one species in particular. And generally, it may be observed
in the writings of most naturalists, that when an organism is described
as intermediate between two _great_ groups, its relations are not to
particular species of either group, but to both groups, as wholes. A
little reflection will show how exceptions (as that of the Lepidosiren,
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