The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
a fish, as does Owen assert that the embryonic jelly-fish is at no time
a polype, but that mammal and fish, jelly-fish and polype pass through
the same state; the mammal and jelly-fish being only further developed
or changed.
{461} _Origin_, Ed. i. p. 439, vi. p. 604.
{462} The uselessness of the branchial arches in mammalia is
insisted on in the _Origin_, Ed. i. p. 440, vi. p. 606. Also the
uselessness of the spots on the young blackbird and the stripes of
the lion-whelp, cases which do not occur in the present Essay.
{463} In the _Origin_, Ed. i. pp. 442, 448, vi. pp. 608, 614 it is
pointed out that in some cases the young form resembles the adult,
_e.g._ in spiders; again, that in the Aphis there is no "worm-like
stage" of development.
{464} In the _Origin_, Ed. i. p. 449, vi. p. 618, the author speaks
doubtfully about the recapitulation theory.
As the embryo, in most cases, possesses a less complicated structure
than that into which it is to be developed, it might have been thought
that the resemblance of the embryo to less complicated forms in the same
great class, was in some manner a necessary preparation for its higher
development; but in fact the embryo, during its growth, may become less,
as well as more, complicated{465}. Thus certain female Epizoic
Crustaceans in their mature state have neither eyes nor any organs of
locomotion; they consist of a mere sack, with a simple apparatus for
digestion and procreation; and when once attached to the body of the
fish, on which they prey, they never move again during their whole
lives: in their embryonic condition, on the other hand, they are
furnished with eyes, and with well articulated limbs, actively swim
about and seek their proper object to become attached to. The larvae,
also, of some moths are as complicated and are more active than the
wingless and limbless females, which never leave their pupa-case, never
feed and never see the daylight.
{465} This corresponds to the _Origin_, Ed. i. p. 441, vi. p. 607,
where, however, the example is taken from the Cirripedes.
_Attempt to explain the facts of embryology._
I think considerable light can be thrown by the theory of descent on
these wonderful embryological facts which are common in a greater or
less degree to the whole animal kingdom, and in some manner to the
vegetable kingdom: on the fact, for instance, of the arteries in the
embryonic mammal, bird, reptile and fish, running and branching in the
same courses and nearly in the same manner with the arteries in the
full-grown fish; on the fact I may add of the high importance to
systematic naturalists{466} of the characters and resemblances in the
embryonic state, in ascertaining the true position in the natural system
of mature organic beings. The following are the considerations which
throw light on these curious points.
{466} _Origin_, Ed. i. p. 449, vi. p. 617.
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