The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
If then the two following propositions are admitted (and I think the
first can hardly be doubted), viz. that variation of structure takes
place at all times of life, though no doubt far less in amount and
seldomer in quite mature life{476} (and then generally taking the form
of disease); and secondly, that these variations tend to reappear at a
corresponding period of life, which seems at least probable, then we
might _a priori_ have expected that in any selected breed the _young_
animal would not partake in a corresponding degree the peculiarities
characterising the _full-grown_ parent; though it would in a lesser
degree. For during the thousand or ten thousand selections of slight
increments in the length of the limbs of individuals necessary to
produce a long-limbed breed, we might expect that such increments would
take place in different individuals (as we do not certainly know at what
period they do take place), some earlier and some later in the embryonic
state, and some during early youth; and these increments would reappear
in their offspring only at corresponding periods. Hence, the entire
length of limb in the new long-limbed breed would only be acquired at
the latest period of life, when that one which was latest of the
thousand primary increments of length supervened. Consequently, the
foetus of the new breed during the earlier part of its existence would
remain much less changed in the proportions of its limbs; and the
earlier the period the less would the change be.
{476} This seems to me to be more strongly stated here than in the
_Origin_, Ed. i.
Public-domain text, read in full here on John Shaqi.
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