The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
A volume might be filled with facts showing what a strong tendency there
is to inheritance, in almost every case of the most trifling, as well as
of the most remarkable congenital peculiarities{191}. The term
congenital peculiarity, I may remark, is a loose expression and can only
mean a peculiarity apparent when the part affected is nearly or fully
developed: in the Second Part, I shall have to discuss at what period of
the embryonic life connatal peculiarities probably first appear; and I
shall then be able to show from some evidence, that at whatever period
of life a new peculiarity first appears, it tends hereditarily to appear
at a corresponding period{192}. Numerous though slight changes, slowly
supervening in animals during mature life (often, though by no means
always, taking the form of disease), are, as stated in the first
paragraphs, very often hereditary. In plants, again, the buds which
assume a different character from their stock likewise tend to transmit
their new peculiarities. There is not sufficient reason to believe that
either mutilations{193} or changes of form produced by mechanical
pressure, even if continued for hundreds of generations, or that any
changes of structure quickly produced by disease, are inherited; it
would appear as if the tissue of the part affected must slowly and
freely grow into the new form, in order to be inheritable. There is a
very great difference in the hereditary tendency of different
peculiarities, and of the same peculiarity, in different individuals and
species; thus twenty thousand seeds of the weeping ash have been sown
and not one come up true;--out of seventeen seeds of the weeping yew,
nearly all came up true. The ill-formed and almost monstrous "Niata"
cattle of S. America and Ancon sheep, both when bred together and when
crossed with other breeds, seem to transmit their peculiarities to their
offspring as truly as the ordinary breeds. I can throw no light on these
differences in the power of hereditary transmission. Breeders believe,
and apparently with good cause, that a peculiarity generally becomes
more firmly implanted after having passed through several generations;
that is if one offspring out of twenty inherits a peculiarity from its
parents, then its descendants will tend to transmit this peculiarity to
a larger proportion than one in twenty; and so on in succeeding
generations. I have said nothing about mental peculiarities being
inheritable for I reserve this subject for a separate chapter.
{191} See _Origin_, Ed. i. p. 13.
{192} _Origin_, Ed. i. p. 86, vi. p. 105.
{193} It is interesting to find that though the author, like his
contemporaries, believed in the inheritance of acquired characters,
he excluded the case of mutilation.
_Causes of Variation._
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