The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
Many practical illustrations of this rapid tendency to increase are on
record, namely during peculiar seasons, in the extraordinary increase of
certain animals, for instance during the years 1826 to 1828, in La
Plata, when from drought, some millions of cattle perished, the whole
country _swarmed_ with innumerable mice: now I think it cannot be
doubted that during the breeding season all the mice (with the exception
of a few males or females in excess) ordinarily pair; and therefore that
this astounding increase during three years must be attributed to a
greater than usual number surviving the first year, and then breeding,
and so on, till the third year, when their numbers were brought down to
their usual limits on the return of wet weather. Where man has
introduced plants and animals into a new country favourable to them,
there are many accounts in how surprisingly few years the whole country
has become stocked with them. This increase would necessarily stop as
soon as the country was fully stocked; and yet we have every reason to
believe from what is known of wild animals that _all_ would pair in the
spring. In the majority of cases it is most difficult to imagine where
the check falls, generally no doubt on the seeds, eggs, and young; but
when we remember how impossible even in mankind (so much better known
than any other animal) it is to infer from repeated casual observations
what the average of life is, or to discover how different the percentage
of deaths to the births in different countries, we ought to feel no
legitimate surprise at not seeing where the check falls in animals and
plants. It should always be remembered that in most cases the checks are
yearly recurrent in a small regular degree, and in an extreme degree
during occasionally unusually cold, hot, dry, or wet years, according to
the constitution of the being in question. Lighten any check in the
smallest degree, and the geometrical power of increase in every
organism will instantly increase the average numbers of the favoured
species. Nature may be compared to a surface, on which rest ten thousand
sharp wedges touching each other and driven inwards by incessant
blows{230}. Fully to realise these views much reflection is requisite;
Malthus on man should be studied; and all such cases as those of the
mice in La Plata, of the cattle and horses when first turned out in S.
America, of the robins by our calculation, &c., should be well
considered: reflect on the enormous multiplying power _inherent and
annually in action_ in all animals; reflect on the countless seeds
scattered by a hundred ingenious contrivances, year after year, over the
whole face of the land; and yet we have every reason to suppose that the
average percentage of every one of the inhabitants of a country will
_ordinarily_ remain constant. Finally, let it be borne in mind that this
average number of individuals (the external conditions remaining the
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