The Foundations of the Origin of Species: Two Essays written in 1842 and 1844Darwin, Charles
Philosophy
The Foundations of the Origin of Species: Two Essays written in 1842 and 1844
Darwin, Charles
Evolution (Biology); Natural selection
Besides this natural means of selection, by which those individuals are
preserved, whether in their egg or seed or in their mature state, which
are best adapted to the place they fill in nature, there is a second
agency at work in most bisexual animals tending to produce the same
effect, namely the struggle of the males for the females. These
struggles are generally decided by the law of battle; but in the case
of birds, apparently, by the charms of their song{235}, by their beauty
or their power of courtship, as in the dancing rock-thrush of Guiana.
Even in the animals which pair there seems to be an excess of males
which would aid in causing a struggle: in the polygamous animals{236},
however, as in deer, oxen, poultry, we might expect there would be
severest struggle: is it not in the polygamous animals that the males
are best formed for mutual war? The most vigorous males, implying
perfect adaptation, must generally gain the victory in their several
contests. This kind of selection, however, is less rigorous than the
other; it does not require the death of the less successful, but gives
to them fewer descendants. This struggle falls, moreover, at a time of
year when food is generally abundant, and perhaps the effect chiefly
produced would be the alteration of sexual characters, and the selection
of individual forms, no way related to their power of obtaining food, or
of defending themselves from their natural enemies, but of fighting one
with another. This natural struggle amongst the males may be compared in
effect, but in a less degree, to that produced by those agriculturalists
who pay less attention to the careful selection of all the young animals
which they breed and more to the occasional use of a choice male{237}.
{235} These two forms of sexual selection are given in _Origin_,
Ed. i. p. 87, vi. p. 107. The Guiana rock-thrush is given as an
example of bloodless competition.
{236} <Note in original.> Seals? Pennant about battles of seals.
{237} In the Linnean paper of July 1, 1858 the final word is
_mate_: but the context shows that it should be _male_; it is
moreover clearly so written in the MS.
_Differences between "Races" and "Species":--first, in their trueness or
variability._
Public-domain text, read in full here on John Shaqi.
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