The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
_Ungulata._--Horses are very interesting. In Europe they date back under
various forms to the Miocene period, and true _Equus_ to the Older
Pliocene. In North America they are chiefly Pliocene, true _Equus_ being
Post-Pliocene, with perhaps one or two species Newer Pliocene; but numerous
ancestral forms date back to the Miocene and Eocene, giving a more perfect
"pedigree of the horse" than the European forms, and going back to a more
primitive type--_Orohippus_. In South America, _Equus_ is the only genus,
and is Post-Pliocene or at most Newer Pliocene. While, therefore, the
ancient progenitors of the Equidæ were common to North America and Europe,
in Miocene and even Eocene times, true horses appear to have arisen in the
Palæarctic region, to have passed into North America in the latter part of
the Pliocene period, and thence to have spread over all suitable districts
in South America. They were not, however, able to maintain themselves
permanently in their new territory, and all became extinct; while in their
birth-place, the Old World, they continue to exist under several varied
forms.
True tapirs are an Old World group. They go back to the Lower Miocene in
Europe, while in both North and South America they are exclusively
Post-Pliocene. They occur in France down to the Newer Pliocene, and must,
about that time, have entered America. The land connection by which this
and so many other animals passed between the Old and New Worlds in late
Tertiary times, was almost certainly in the North Pacific, south of
Behring's Straits, where, as will be seen by our general map, there is a
large expanse of shallow water, which a moderate elevation would convert
into dry land, in a sufficiently temperate latitude.
{155}The peccary (_Dicotyles_), now a characteristic South American genus,
is a recent immigrant from North America, where it appears to have been
developed from ancestral forms of swine dating back to the Miocene period.
Antelopes are an Old World type, but a few of them appear to have entered
North, and reached South America in late Pliocene times. Camels, strange to
say, are a special North American type, since they abounded in that
continent under various ancient forms in the Miocene period. Towards the
end of that period they appear to have entered eastern Asia, and developed
into the Siberian _Merycotherium_ and the North Indian _Camelus_, while in
the Pliocene age the ancestral llamas entered South America.
_Cervidæ_ are a wide-spread northern type in their generalized form, but
true deer (_Cervus_) are Palæarctic. They abounded in Europe in Miocene
times, but only appear in North and South America in the later Pliocene and
Post-Pliocene periods.
True oxen (_Bovinæ_) seem to be an Oriental type (Miocene), while they
appear in Europe only late in the Pliocene period, and in America are
confined to the Post-Pliocene.
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