The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
Of Carabidæ Mr. Bates enumerates 244 species belonging to 84 genera, and by
comparing these with the Coleoptera of a tract of about equal extent in
western Europe, he concludes that there is little similarity, and that the
cases of affinity to the forms of eastern tropical Asia preponderate. By
comparing his genera with the distributions as given in _Gemminger and
Harold's Catalogue_, a somewhat different result is arrived at. Leaving out
the generic types altogether peculiar to Japan, and also those genera of
such world-wide distribution that they afford no clear indications for our
purpose, it appears that no less than twenty-two genera, containing
seventy-four of the Japanese species, are either exclusively Palæarctic,
Palæarctic and Nearctic, or highly characteristic of the Palæarctic region;
then come thirteen genera containing eighty-seven of the species which have
a very wide distribution, but are also Palæarctic: we next have seventeen
genera containing twenty-four of the Japanese species which are decidedly
Oriental and tropical. Here then the fair comparison is between the
twenty-two genera and seventy-four species whose affinities are clearly
Palæarctic or at least north temperate, and seventeen genera with
twenty-four species which are Asiatic and tropical; and this seems to prove
that, although South {229}Japan (like North China) has a considerable
infusion of tropical forms, there is a preponderating substratum of
Palæarctic forms, which clearly indicate the true position of the islands
in zoological geography. There are also a few cases of what may be called
eccentric distribution; which show that Japan, like many other
island-groups, has served as a kind of refuge in which dying-out forms
continue to maintain themselves. These, which are worthy of notice, are as
follows: _Orthotrichus_ (1 sp.) has the only other species in Egypt;
_Trechichus_ (1 sp.) has two other species, of which one inhabits Madeira,
the other the Southern United States; _Perileptus_ (1 sp.) has two other
species, of which one inhabits Bourbon, the other West Europe; and lastly,
_Crepidogaster_ (1 sp.) has the other known species in South Africa. These
cases diminish the value of the indications afforded by some of the
Japanese forms, whose only allies are single species in various remote
parts of the Oriental region.
The Staphylinidæ have been described by Dr. Sharp, and his list exhibits a
great preponderance of north temperate, or cosmopolitan forms, with a few
which are decidedly tropical. The Pselaphidæ and Scydmenidæ, also described
by Dr. Sharp, exhibit, according to that gentleman, "even a greater
resemblance to those of North America than to those of Europe," but he says
nothing of any tropical affinities. The water-beetles are all either
Palæarctic or of wide distribution.
The Lucanidæ (_Gemm. and Har. Cat._, 1868) exhibit an intermingling of
Palæarctic and Oriental genera.
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