The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
{65}We see, then, that even the exceedingly rich and isolated Neotropical
region is less rich and diversified in its forms of mammalian life than the
very much smaller area of the Indian region, or the temperate Palæarctic,
and very much less so than the Ethiopian region; while even the
comparatively poor Nearctic region, is nearly equal to it in the number of
its family types. If these were united they would possess fifty-five
families, a number very disproportionate to those of the remaining two.
Another consideration is, that although the absence of certain forms of
life makes a region more isolated, it does not make it zoologically more
important; for we have only to suppose some five or six families, now
common to both, to become extinct either in the Ethiopian or the Indian
regions, and they would become as strongly differentiated from all other
regions as South America, while still remaining as rich in family types. In
birds exactly the same phenomenon recurs, the family types being less
numerous in South America than in either of the other tropical regions of
the earth, but a larger proportion of them are restricted to it. It will be
shown further on, that the Ethiopian and Indian, (or, as I propose to call
it in this work, Oriental) regions, are sufficiently differentiated by very
important groups of animals peculiar to each; and that, on strict
zoological principles they are entitled to rank as regions of equal value
with the Neotropical and Australian. It is perhaps less clear whether the
Palæarctic should be separated from the Oriental region, with which it has
undoubtedly much in common; but there are many and powerful reasons for
keeping it distinct. There is an unmistakably different facies in the
animal forms of the two regions; and although no families of mammalia or
birds, and not many genera, are wholly confined to the Palæarctic region, a
very considerable number of both have their metropolis in it, and are very
richly represented. The distinction between the characteristic forms of
life in tropical and cold countries is, on the whole, very strongly marked
in the northern hemisphere; and to refuse to recognise this in a
subdivision of the earth which is established for the very purpose of
expressing such contrasts more clearly and concisely than by ordinary
geographical terminology, would be both illogical and {66}inconvenient. The
one question then remains, whether the Nearctic region should be kept
separate, or whether it should form part of the Palæarctic or of the
Neotropical regions. Professor Huxley and Mr. Blyth advocate the former
course; Mr. Andrew Murray (for mammalia) and Professor Newton (for birds)
think the latter would be more natural. No doubt much is to be said for
both views, but both cannot be right; and it will be shown in the latter
part of this chapter that the Nearctic region is, on the whole, fully as
well defined as the Palæarctic, by positive characters which differentiate
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