The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
_Objections to the system of Circumpolar Zones._--Mr. Allen's system of
"realms" founded on climatic zones (given at p. 61), having recently
appeared in an ornithological work of considerable detail and research,
calls for a few remarks. The author continually refers to the "_law of the
distribution of life in circumpolar zones_," as if it were one generally
accepted and that admits of no dispute. But this supposed "law" only
applies to the smallest details of distribution--to the range and
increasing or decreasing numbers of _species_ as we pass from north to
south, or the reverse; while it has little bearing on the great features of
zoological geography--the limitation of groups of _genera_ and _families_
to certain areas. It is analogous to the "_law of adaptation_" in the
organisation of animals, by which members of various groups are suited for
an aerial, an aquatic, a desert, or an arboreal life; are herbivorous,
carnivorous, or insectivorous; are fitted to live underground, or in fresh
waters, or on polar ice. It was once thought that these adaptive
peculiarities were suitable foundations for a classification,--that whales
were fishes, and bats birds; and even to this day there are naturalists who
cannot recognise the essential diversity of structure in such groups as
swifts and swallows, sun-birds and humming-birds, under the superficial
disguise caused by adaptation to a similar mode of life. The application of
Mr. Allen's principle leads to equally erroneous results, as may be well
seen by considering his separation of "the southern third of Australia" to
unite it with New Zealand as one of his secondary zoological divisions. If
there is one country in the world whose fauna is strictly homogeneous, that
country is Australia; while New Guinea on the one hand, and New Zealand on
the other, are as sharply differentiated from Australia as any adjacent
parts of the same primary zoological division can possibly be. Yet the
"_law of circumpolar distribution_" leads to the division of {68}Australia
by an arbitrary east and west line, and a union of the northern two-thirds
with New Guinea, the southern third with New Zealand. Hardly less unnatural
is the supposed equivalence of South Africa (the African temperate realm)
to all tropical Africa and Asia, including Madagascar (the Indo-African
tropical realm). South Africa has, it is true, some striking peculiarities;
but they are absolutely unimportant as compared with the great and radical
differences between tropical Africa and tropical Asia. On these examples we
may fairly rest our rejection of Mr. Allen's scheme.
Public-domain text, read in full here on John Shaqi.
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