The Geographical Distribution of Animals, Volume 2: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 2: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
The most striking fact presented by this order, from our present point of
view, is the strict limitation of well-marked families to definite areas.
The Cebidæ and Hapalidæ would alone serve to mark out tropical America as
the nucleus of one of the great zoological divisions of the earth. In the
Eastern Hemisphere, the corresponding fact is the entire absence of the
order from the Australian region, with the exception of one or two outlying
forms, which have evidently transgressed the normal limits of their group.
The separation of the Ethiopian and Oriental regions is, in this order,
mainly indicated by the distribution of the genera, no one of which is
common to the two regions. The two highest families, the Simiidæ and the
Semnopithecidæ, are pretty equally distributed about two equatorial foci,
one situated in West Africa, the other in the Malay archipelago,--in Borneo
or the Peninsula of Malacca;--while the third family, Cynopithecidæ, ranges
over the whole of both regions, and somewhat overpasses their limits. The
Lemuroid group, on the other hand, offers us one of the most singular
phenomena in geographical distribution. It consists of three families, the
species of which are grouped into six sub-families and 13 genera. One of
these families and two of the sub-families, comprising 7 genera, and no
less than 30 out of the total of 50 species, are confined to the one island
of Madagascar. Of the remainder, 3 genera, comprising 15 species, are
spread over tropical Africa; while three other genera with 5 species,
inhabit certain restricted portions of India and the Malay islands. These
curious facts point unmistakably to the former existence of a large tract
of land in what is now the Indian Ocean, connecting Madagascar on the one
hand with Ceylon, and with the Malay countries on the {180}other. About
this same time (but perhaps not contemporaneously) Madagascar must have
been connected with some portion of Southern Africa, and the whole of the
country would possess no other Primates but Lemuroidea. After the
Madagascar territory (very much larger than the existing island) had been
separated, a connection appears to have been long maintained (probably by a
northerly route) between the more equatorial portions of Asia and Africa;
till those higher forms had become developed, which were afterwards
differentiated into _Simia_, _Presbytes_, and _Cynopithecus_, on the one
hand, and into _Troglodytes_, _Colobus_, and _Cynocephalus_, on the other.
In accordance with the principle of competition so well expounded by Mr.
Darwin, we can understand how, in the vast Asiatic and African area north
of the Equator, with a great variety of physical conditions and the
influence of a host of competing forms of life, higher types were developed
than in the less extensive and long-isolated countries south of the
Equator. In Madagascar, where these less complex conditions prevailed in a
considerable land-area, the lowly organized Lemuroids have diverged into
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