The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
Witham in 1833 described several Carboniferous species of pine-wood,
under the generic name Pinites, separating under the name _Pitus_ species
which appeared to have the discs on the cell-walls separate and in
transverse lines. Witham's name was changed by Goeppert to _Araucarites_,
to indicate the similarity of these woods to Araucaria, _Pinites_ being
reserved for trees more closely allied to the ordinary pines. Endlicher,
restricting Araucarites to foliage, etc., of Araucaria-like trees, gave
the name _Dadoxylon_ to the wood; and this, through Unger's "Genera and
Species," has gained somewhat general acceptance. Endlicher also gave the
name _Pissadendron_ to the species which Witham had called _Pitus_; but
Brongniart proposed the name _Palæoxylon_ to include all the species with
thick and complex medullary rays, whatever the arrangement of the discs.
In Schimper's new work Kraus substitutes _Araucarioxylon_ for Endlicher's
_Dadoxylon_, and includes under _Pissadendron_ all the species placed by
Brongniart in _Palæoxylon_.
To understand all this confusion, it may be observed that the characters
available in the determination of Palæozoic coniferous wood are chiefly
the form and arrangement of the wood-cells, the character of the bordered
pores or discs of their walls, and the form and composition of the
medullary rays.
The character on which Witham separated his genus _Pitus_ from _Pinites_
is, as I have ascertained by examination of slices of one of his original
specimens kindly presented to me by Mr. Sanderson, of Edinburgh,
dependent on state of preservation, the imperfectly preserved discs
or areolations of the walls of the fibre presenting the appearance of
separate and distinct circles, while in other parts of the same specimens
these discs are seen to be contiguous and to assume hexagonal forms, so
that in this respect they do not really differ from the ordinary species
of _Dadoxylon_. The true character for subdividing those species which
are especially characteristic of the Carboniferous, is the composite
structure of the medullary rays, which are thick and composed of several
radial piles of cells placed side by side. This was the character
employed by Brongniart in separating the genus _Palæoxylon_, though he
might with convenience have retained Witham's name, merely transferring
to the genus the species of Witham's _Pinites_ which have complex
medullary rays. The Erian rocks present the greatest variety of types,
and _Palæoxylon_ is especially characteristic of the Lower Carboniferous,
while species of _Dadoxylon_ with two rows of bordered pores and simple
medullary rays are especially plentiful in the upper coal-formation and
Permo-Carboniferous.
The following table will clearly show the distinctive characters and
relations of the genera in question, as held by the several authors above
referred to:
_Wood of Palæozoic Conifers._
Public-domain text, read in full here on John Shaqi.
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