The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
It will be observed that I regard the striated and ribbed stems not as
internal axes, but as representing the outer surface of the plants. This
was certainly the case with the present species and with _C. Suckovii_
and _C. nodosus_. Other species, and especially those which belonged to
Calamodendron, no doubt had a smooth or irregularly wrinkled external
bark; but this gives no good ground for the manner in which some writers
on this subject confound Calamites with Calamodendra, and both with
Asterophyllites and Sphenophyllum. With this no one who has studied these
plants, rooted in their native soils, and with their appendages still
attached, can for a moment sympathise. One of the earliest geological
studies of the writer was a bed of these erect Calamites, which he
showed to Sir C. Lyell in 1844, and described in the "Proceedings of
the Geological Society" in 1851, illustrating the habit of growth as
actually seen well exposed in a sandstone cliff. Abundant opportunities
of verifying the conclusions formed at that time have since occurred,
the results of which have been summed up in the figures in Acadian
Geology, which, though they have been treated by some botanists as merely
restorations, are in reality representations of facts actually observed.
On these subjects, without entering into details, and referring for these
to the elaborate discussions of Schimper, Williamson, and McNab, and to
my paper on the subject, "Journal of the Geological Society," vol. xxvii,
p. 54, I may remark:
1. That the aërial stems of ordinary Calamites had a thin cortical layer,
with lacunæ and fibrous bundles and multiporous vessels--the whole not
differing much from the structure of modern Equiseta.
2. Certain arborescent forms, perhaps allied to the true Calamites,
as well as possibly the old underground stems of ordinary species[DC]
assumed a thick-walled character in which the tissues resembled the
wedges of an exogen, and abundance of pseudo-scalariform fibres were
developed, while the ribbing of the external surface became obsolete or
was replaced by a mere irregular wrinkling.
[DC] Williamson, "Transactions of the Royal Society." McNab, in
"Proceedings of the Edinburgh Botanical Society."
3. Sufficient discrimination has not been exercised in separating casts
of the internal cavities of Calamites and Calamodendron from those
representing other surfaces and the proper external surface.
4. There is no excuse for attributing to Calamites the foliage of
Annularia, Asterophyllites, and Sphenophyllum, since these leaves have
not been found attached to true Calamite stems, and since the structure
of the stems of Asterophyllites as described by Williamson, and that of
Sphenophyllum as described by the writer,[DD] are essentially different
from those of Calamites.
[DD] "Journal of the Geological Society," 1866.
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