The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
Under this name we may provisionally include those rounded spherical
bodies found in the coal and its accompanying beds, and also in
the Erian, which may be regarded as Macrospores or Sporocarps of
Protosalvinia, or other Rhizocarpean plants akin to those described above
in Chapter III, which see for description.
_Genus_ Protosalvinia.--Under this we include sporocarps allied to those
of _Salvinia_, as described in Chapter III.
_Family_ Filices.
Under this head I shall merely refer to a few groups of special interest,
and to the provisional arrangement adopted for the fronds of ferns when
destitute of fructification.
The external appearances of trunks of tree-ferns have been already
referred to.
With respect to tree ferns, the oldest known examples are those from
the Middle Devonian of New York and Ohio, which I have described in the
"Journal of the Geological Society," 1871 and 1881. As these are of some
interest, I have reproduced their descriptions in a note appended to
Chapter III, which see.
The other forms most frequently occurring in the Carboniferous are
_Caulopteris_, _Palæopteris_, and _Megaphyton_[DE] Stems showing merely
masses of aërial roots are known by the name _Psaronius_.
[DE] See my "Acadian Geology," also below.
With reference to the classification of Palæozoic ferns, this has
hitherto been quite arbitrary, being based on mere form and venation of
fronds, but much advance has recently been made in the knowledge of their
fructification, warranting a more definite attempt at classification. The
following are provisional genera usually adopted:
1. _Cyclopteris_, Brongniart.--Leaflets more or less rounded or
wedge-shaped, without midrib, the nerves spreading from the point of
attachment. This group includes a great variety of fronds evidently of
different genera, were their fructification known; and some of them
probably portions of fronds, the other parts of which may be in the next
genus.
2. _Neuropteris_, Brongniart.--Fronds pinnate, and with the leaflets
narrowed at the base; midrib often not distinct, and disappearing toward
the apex. Nervures equal, and rising at an acute angle. Ferns of this
type are among the most abundant in the coal-formation.
3. _Odontopteris_, Brongniart.--In these the frond is pinnate, and
the leaflets are attached by their whole base, with the nerves either
proceeding wholly from the base, or in part from an indistinct midrib,
which soon divides into nervures.
4. _Dictyopteris_, Gutbier.--This is a beautiful style of fern, with
leaflets resembling those of _Neuropteris_, but the veins arranged
in a network of oval spaces. Only a few species are known in the
coal-formation.
5. _Lonchopteris_, Brongniart.--Ferns with netted veins like the above,
but with a distinct midrib, and the leaflets attached by the whole base.
Of this, also, we can boast but few species.
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