The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
This stem is probably the upper part of one or other of the species
of _Psaronius_ found in the same bed (_P. Erianus_, Dawson, and _P.
textilis_, Dawson).[BG] It appears to have been an erect stem embedded
in situ in sandstone, and preserved as a cast. The stem is small, being
only two inches, or a little more, in diameter. It is coarsely wrinkled
longitudinally, and covered with large leaf-scars, each an inch in
diameter, of a horseshoe-shape. The petioles, five of which remain,
separate from these scars with a distinct articulation, except at one
point near the base, where probably a bundle or bundles of vessels passed
into the petiole. They retain their form at the attachment to the stem,
but a little distance from it they are flattened. They are inflated
at the base, and somewhat rapidly diminish in size. The leaf-scars
vary in form, and are not very distinct, but they appear to present
a semicircular row of pits above, largest in the middle. From these
there proceed downward a series of irregular furrows, converging to a
second and more obscure semicircle of pits, within or below which is the
irregular scar or break above referred to. The attitude and form of the
petioles will be seen from Fig. 24, _supra_.
[BG] Memoir on Devonian Flora, "Proceedings of the Royal Society," May,
1870.
The petioles are broken off within a few inches of the stem; but other
fragments found in the same beds appear to show their continuation, and
some remains of their foliage. One specimen shows a series of processes
at the sides, which seem to be the remains of small pinnæ, or possibly
of spines on the margin of the petiole. Other fragments show the
division of the frond, at first in a pinnate manner, and subsequently by
bifurcation; and some fragments show remains of pinnules, possibly of
fertile pinnules. These are very indistinct, but would seem to show that
the plant approached, in the form of its fronds and the arrangement of
its fructification, to the Cyclopterids of the sub-genus _Aneimites_,
one of which (_Aneimites Acadica_), from the Lower Carboniferous of Nova
Scotia, I have elsewhere described as probably a tree-fern,[BH] The
fronds were evidently different from those of _Archæopteris_[BI] a genus
characteristic of the same beds, but of very different habit of growth.
This accords with the fact that there is in Prof. Hall's collection a
mass of fronds of _Cyclopteris_ (_Archæopteris_) Jacksoni, so arranged
as to make it probable that the plant was an herbaceous fern, producing
tufts of fronds on short stems in the ordinary way. The obscurity of
the leaf-scars may render it doubtful whether the plant above described
should be placed in the genus _Caulopteris_ or in _Stemmatopteris_; but
it appears most nearly allied to the former. The genus is at present,
of course, a provisional one; but I have thought it only justice to the
diligent labours of Mr. Lockwood to name this curious and interesting
fossil _Caulopteris Lockwoodi_.
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