The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
I have already stated that the Carboniferous constitutes the headquarters
of the _Cordaites_ (Fig. 58), of which a large number of species have
been described, both in Europe and America. We sometimes, though
rarely, find their stems showing structure. In this case we have a
large cellular pith, often divided by horizontal partitions into flat
chambers, and constituting the objects which, when detached, are called
_Sternbergiæ_ (Fig. 62). These Sternbergia piths, however, occur in true
conifers as well, as they do in the modern world in some trees, like our
common butternut, of higher type; and I showed many years ago that the
Sternbergia type may be detected in the young twigs of the balsam-fir
(_Abies balsamifera_). The pith was surrounded by a ring of scalariform
or barred tissue, often of considerable thickness, and in young stems so
important as to have suggested lycopodiaceous affinities. But as the stem
grew in size, a regular ring of woody wedges, with tissue having rounded
or hexagonal pores or discs, like those of pines, was developed. Outside
this was a bark, often apparently of some thickness. This structure in
many important points resembles that of cycads, and also approaches to
the structure of Sigillaria, while in its more highly developed forms it
approximates to that of the conifers.
[Illustration: Fig. 58.--_Cordaites_ (_Dorycordaites_), Grand d'Eury,
reduced.]
[Illustration: Fig. 59.--Fruits of _Cordaites_ and Taxine Conifers
(coal-formation. Nova Scotia.) A, _Antholithes squamosus_ (two thirds).
B, _A. rhabdocarpi_. (two thirds). B^1, Carpel restored. C, _A. spinosus_
(natural size). D, _Trigonocarpum intermedium_. E, _T. Noeggerathii_.
F, _T. avellanum_. G, _Rhabdocarpus insignis_, reduced. H, _Antholithes
pygmæus_. I, _Cardiocarpum fluitans_. K, _Cardiocarpum bisectum_. L,
_Sporangites papillata_, lycopodiaceous macrospores (natural size and
magnified).]
On the stems so constructed were placed long and often broad many-nerved
leaves, with rows of stomata or breathing-pores, and attached by somewhat
broad bases to the stem and branches. The fruit consisted of racemes, or
clusters of nutlets, which seem to have been provided with broad lateral
wings for flotation in the air, or in some cases with a pulpy envelope,
which flattens into a film. There seem to have been structures of both
these kinds, though in the state of preservation of these curious seeds
it is extremely difficult to distinguish them. In the first case they
must have been intended for dissemination by the wind, like the seeds
of spruces. In the latter case they may have been disseminated like the
fruits of taxine trees by the agency of animals, though what these were
it would be difficult to guess. These trees had very great reproductive
power, since they produced numerous seeds, not singly or a few together,
as in modern yews, but in long spikes or catkins bearing many seeds (Fig.
59).
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