The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
A remarkable fact in connection with them, and showing also the manner in
which the most durable vegetable structures may perish by decay, is that,
like the Cordaites, they had large piths with transverse partitions, a
structure which, as I have already mentioned, appears on a minute scale
in the twigs of the fir-tree, and that sometimes casts of these piths in
sandstone appear in a separate form, constituting what have been named
_Sternbergiæ_ or _Artisiæ_. As Renault well remarks with reference to
Cordaites, the existence of this chambered form of pith implies rapid
elongation of the stem, so that the Cordaites and conifers of the
coal-formation were probably quickly growing trees (Fig. 62).
[Illustration: Fig. 62.--_Sternbergia_ pith of _Dadoxylon_. A, Specimen
(natural size), showing remains of wood at _a_, _a_. B, Junction of
wood and pith, magnified. C, Cells of the wood of do., _a_, _a_; _b_,
medullary ray; _c_, areolation.]
The same general statements may be made as to the coal-vegetation as in
relation to that of the Erian. In the coal period we have found none of
the higher exogens, and there are only obscure and uncertain indications
of the presence of endogens, which we may reserve for a future chapter;
but gymnosperms abound and are highly characteristic. On the other hand,
we have no mosses or lichens, and very few Algæ, but a great number of
ferns and Lycopodiaceæ or club-mosses (Fig. 63). Thus, the coal-formation
period is botanically a meeting-place of the lower phænogams and the
higher cryptogams, and presents many forms which, when imperfectly known,
have puzzled botanists in regard to their position in one or other
series. In the present world, the flora most akin to that of the coal
period is that of warm, temperate regions in the southern hemisphere. It
is not properly a tropical flora, nor is it the flora of a cold region,
but rather indicative of a moist and equable climate. Still, we must bear
in mind that we may often be mistaken in reasoning as to the temperature
required by extinct species of plants, differing from those now in
existence. Further, we must not assume that the climatal conditions of
the northern hemisphere were in the coal period at all similar to those
which now prevail. As Sir Charles Lyell has shown, a less amount of land
in the higher latitudes would greatly modify climates, and there is
every reason to believe that in the coal period there was less land than
now. Further, it has been shown by Tyndall that a very small additional
amount of carbonic acid in the atmosphere would, by obstructing the
radiation of heat from the earth, produce almost the effect of a glass
roof or conservatory, extending over the whole world. Again, there is
much in the structure of the leaves of the coal-plants, as well as in
the vast amount of carbon which they accumulated in the form of coal,
and the characteristics of the animal life of the period, to indicate,
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