Some animals have the right ventricle smooth internally, but the
left provided with fibrous bands, such as the goose, swan, and
larger birds; and the reason is the same here as elsewhere. As
the lungs are spongy and loose and soft, no great amount of force
is required to force the blood through them; therefore the right
ventricle is either without the bundles in question, or they are
fewer and weaker, and not so fleshy or like muscles. Those of the
left ventricle, however, are both stronger and more numerous,
more fleshy and muscular, because the left ventricle requires to
be stronger, inasmuch as the blood which it propels has to be
driven through the whole body. And this, too, is the reason why
the left ventricle occupies the middle of the heart, and has
parietes three times thicker and stronger than those of the right
Hence all animals--and among men it is similar--that are endowed
with particularly strong frames, and with large and fleshy limbs
at a great distance from the heart, have this central organ of
greater thickness, strength, and muscularity. This is manifest
and necessary. Those, on the contrary, that are of softer and
more slender make have the heart more flaccid, softer, and
internally either less or not at all fibrous. Consider, farther,
the use of the several valves, which are all so arranged that the
blood, once received into the ventricles of the heart, shall
never regurgitate; once forced into the pulmonary artery and
aorta, shall not flow back upon the ventricles. When the valves
are raised and brought together, they form a three-cornered line,
such as is left by the bite of a leech; and the more they are
forced, the more firmly do they oppose the passage of the blood.
The tricuspid valves are placed, like gate-keepers, at the
entrance into the ventricles from the venae cavae and pulmonary
veins, lest the blood when most forcibly impelled should flow
back. It is for this reason that they are not found in all
animals, nor do they appear to have been constructed with equal
care, in all animals in which they are found. In some they are
more accurately fitted, in others more remissly or carelessly
contrived, and always with a view to their being closed under a
greater or a slighter force of the ventricle. In the left
ventricle, therefore, in order that the occlusion may be the more
perfect against the greater impulse, there are only two valves,
like a mitre, and produced into an elongated cone, so that they
come together and touch to their middle; a circumstance which
perhaps led Aristotle into the error of supposing this ventricle
to be double, the division taking place transversely. For the
same reason, and that the blood may not regurgitate upon the
pulmonary veins, and thus the force of the ventricle in
propelling the blood through the system at large come to be
neutralized, it is that these mitral valves excel those of the
right ventricle in size and strength and exactness of closing.
Public-domain text, read in full here on John Shaqi.
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