alcoholic ferments have a mode of life which is probably quite
exceptional, since it is not generally met with in other species,
vegetable or animal.
Another equally exceptional characteristic of yeast and
fermentation in general consists in the small proportion which
the yeast that forms bears to the sugar that decomposes. In all
other known beings the weight of nutritive matter assimilated
corresponds with the weight of food used up, any difference that
may exist being comparatively small. The life of yeast is
entirely different. For a certain weight of yeast formed, we may
have ten times, twenty times, a hundred times as much sugar, or
even more decomposed, as we shall experimentally prove by-and-
bye; that is to say, that whilst the proportion varies in a
precise manner, according to conditions which we shall have
occasion to specify, it is also greatly out of proportion to the
weight of the yeast. We repeat, the life of no other being, under
its normal physiological conditions, can show anything similar.
The alcoholic ferments, therefore, present themselves to us as
plants which possess at least two singular properties: they can
live without air, that is without oxygen, and they can cause
decomposition to an amount which, though variable, yet, as
estimated by weight of product formed, is out of all proportion
to the weight of their own substance. These are facts of so great
importance, and so intimately connected with the theory of
fermentation, that it is indispensable to endeavour to establish
them experimentally, with all the exactness of which they will
admit.
The question before us is whether yeast is in reality an
anaerobian [Footnote: Capable of living without free oxygen--a
term invented by Pasteur.--En.] plant, and what quantities of
sugar it may cause to ferment, under the various conditions under
which we cause it to act.
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