estimate of their power of decomposition, we may be led to
conclude that in the first case that power was entirely wanting,
and that in the second case it was considerable, although all the
time we are dealing with the same organism--the identical
ferment.
M. Schutzenberger is astonished that fermentation can take place
in the presence of free oxygen, if, as we suppose, the
decomposition of the sugar is the consequence of the nutrition of
the yeast, at the expense of the combined oxygen, which yields
itself to the ferment. At all events, he argues, fermentation
ought to be slower in the presence of free oxygen. But why should
it be slower? We have proved that in the presence of oxygen the
vital activity of the cells increases, so that, as far as
rapidity of action is concerned, its power cannot be diminished.
It might, nevertheless, be weakened as a ferment, and this is
precisely what happens. Free oxygen imparts to the yeast a vital
activity, but at the same time impairs its power as yeast--qua
yeast, inasmuch as under this condition it approaches the state
in which it can carry on its vital processes after the manner of
an ordinary fungus; the mode of life, that is, in which the ratio
between the weight of sugar decomposed and the weight of the new
cells produced will be the same as holds generally among
organisms which are not ferments. In short, varying our form of
expression a little, we may conclude with perfect truth, from the
sum total of observed facts, that the yeast which lives in the
presence of oxygen and can assimilate as much of that gas as is
necessary to its perfect nutrition, ceases absolutely to be a
ferment at all. Nevertheless, yeast formed under these conditions
and subsequently brought into the presence of sugar, OUT OF THE
INFLUENCE OF AIR, would decompose more IN A GIVEN TIME than in
any other of its states. The reason is that yeast which has
formed in contact with air, having the maximum of free oxygen
that it can assimilate is fresher and possessed of greater vital
activity than that which has been formed without air or with an
insufficiency of air. M. Schutzenberger would associate this
activity with the notion of time in estimating the power of the
ferment; but he forgets to notice that yeast can only manifest
this maximum of energy under a radical change of its life
conditions; by having no more air at its disposal and breathing
no more free oxygen. In other words, when its respiratory power
becomes null, its fermentative power is at its greatest. M.
Schutzenberger asserts exactly the opposite (p. 151 of his work--
Paris, 1875) [Footnote: Page 182, English edition], and so
gratuitously places himself in opposition to facts.
Public-domain text, read in full here on John Shaqi.
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