The History of Creation, Vol. 2 (of 2): Or the Development of the Earth and its Inhabitants by the Action of Natural CausesHaeckel, Ernst
Science
The History of Creation, Vol. 2 (of 2): Or the Development of the Earth and its Inhabitants by the Action of Natural Causes
Haeckel, Ernst
Evolution; Heredity; Human beings -- Origin
Hence the Star-fishes would be compound stocks of worms which, by the
radial formation of buds, have developed out of true articulated worms,
or Annelids. This hypothesis is most strongly supported by the
comparative anatomy, and by the ontogeny of some Star-fishes (Colastra),
and of segmented worms. The many-jointed Ring-worms (Annelida) in their
inner structure are closely allied to the individual arms or radii of
the Star-fishes, that is to the original single worms, which each arm
represents. Each of the five worms of the Star-fish is a chain composed
of a great number of equi-formal members, or metamera, lying one behind
the other, like every segmented Worm, and every Arthropod. As in the
latter a central nervous cord, the ventral nerve cord runs along the
central line of the ventral wall of each segment. On each metameron
there is a pair of non-jointed feet, and besides these, in most cases,
one or more hard thorns or bristles similar to those of many Ring-worms.
A detached arm of a Star-fish can lead an independent life, and can
then, by the radially-directed growth of buds at one end, again become a
complete star.
The most important proofs, however, of the truth of my hypothesis are
furnished by the ontogeny or the individual development of the
Echinoderma. The most remarkable facts of this ontogeny were first
discovered in the year 1848 by the great zoologist, Johannes Müller of
Berlin. Some of its most important stages are represented on Plates
VIII. and IX. (Compare their explanation in the Appendix.) Fig. _A_ on
Plate IX. shows us a common Sea-star (Uraster), Fig. _B_, a Sea-lily
(Comatula), Fig. _C_, a Sea-urchin (Echinus), and Fig. _D_, a
Sea-cucumber (Synapta). In spite of the extraordinary difference of form
manifested by these four representatives of the different classes of
Star-fishes, yet the beginning of their development is identical in all
cases. Out of the egg an animal-form develops which is utterly
different from the fully developed Star-fish, but very like the ciliated
larvæ of certain segmented Worms (Star-worms and Ring-worms). This
peculiar animal-form is generally called the “larva,” but more correctly
the “nurse” of these Star-fish. It is very small and transparent, swims
about by means of a fringe of cilia, and is always composed of two equal
symmetrical halves or sides. The fully grown Echinoderm, however—which
is frequently more than a hundred times larger, and quite opaque—creeps
at the bottom of the sea, and is always composed of at least five
co-ordinate pieces, or antimera, in the form of radii. Plate VIII. shows
the development of the “nurses” of the four Echinoderms represented on
Plate IX.
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